Osteology and phylogeny ofRobustichthys luopingensis, the largest holostean fish in the Middle Triassic

Allis, 1909, The cranial anatomy of the mail-cheeked fishes, Zoologica, 22, 1

Alvarado-Ortega, 2008, A new genus of ionoscopiform fish (Halecomorphi) from the Lower Cretaceous (Albian) lithographic limestones of the Tlayúa quarry, Puebla, México, Journal of Paleontology, 82, 163, 10.1666/04-152.1

Applegate, 1988, A new genus and species of a holostean belonging to the family Ophiopsidae, Teoichthys kallistos, from the Cretaceous near Tepexi de Rodríguez, Puebla, Instituto de Geología, Revista, 7, 200

Arratia, 1999, The monophyly of Teleostei and stem-group teleosts. Consensus and disagreements, Mesozoic Fishes 2—Systematics and Fossil Record, 265

Arratia, 2013, Morphology, taxonomy, and phylogeny of Triassic pholidophorid fishes (Actinopterygii, Teleostei), Journal of Vertebrate Paleontology, 33, 1, 10.1080/02724634.2013.835642

Arratia, 1991, Palatoquadrate and its ossifications: development and homology within osteichthyans, Journal of Morphology, 208, 1, 10.1002/jmor.1052080102

Azuma, 2008, Mitogenomic evaluation of the historical biogeography of cichlids toward reliable dating of teleostean divergences, BMC Evolutionary Biology, 8, 215, 10.1186/1471-2148-8-215

Bartram, 1975, The holostean fish genus Ophiopsis Agassiz, Zoological Journal of the Linnean Society, 56, 183, 10.1111/j.1096-3642.1975.tb00263.x

Bartram, 1977, The Macrosemiidae, a Mesozoic family of holostean fishes. Bulletin of the British Museum, natural history, Geology, 29, 137

Brito, 1988, La structure du suspensorium de Vinctifer, poisson actinoptérygien mésozoïque: Remarques sur les implications phylogénétiques, Geobios, 21, 819, 10.1016/s0016-6995(88)80098-6

Brito, 2013, Cipactlichthys scutatus, gen. nov., sp. nov. a new halecomorph (Neopterygii, Holostei) from the Lower Cretaceous Tlayúa Formation of Mexico, PLOS ONE, 8, e73551, 10.1371/journal.pone.0073551

Broughton, 2013, Multi-locus phylogenetic analysis reveals the pattern and tempo of bony fish evolution, PLOS Currents Tree of Life, 10.1371/currents.tol.2ca8041495ffafd0c92756e75247483e

Cavin, 2010, Diversity of Mesozoic semionotiform fishes and the origin of gars (Lepisosteidae), Naturwissenschaften, 97, 1035, 10.1007/s00114-010-0722-7

Cavin, 2013, Osteology and relationships of Thaiichthys nov. gen.: a Ginglymodi from the Late Jurassic–Early Cretaceous of Thailand, Palaeontology, 56, 183, 10.1111/j.1475-4983.2012.01184.x

Cavin, 2006, A new Semionotiform (Actinopterygii, Neopterygii) from Upper Jurassic–Lower Cretaceous deposits of North-East Thailand, with comments on the relationships of Semionotiforms, Palaeontology, 49, 339, 10.1111/j.1475-4983.2006.00539.x

Chen, 2014, A new species of Sangiorgioichthys Tintori & Lombardo, 2007 (Actinopterygii; Semionotiformes) from the Pelsonian (Anisian, Middle Triassic) of Guizhou Province, South China, Neues Jahrbuch für Geologie und Paläontologie—Abhandlungen, 273, 65, 10.1127/0077-7749/2014/0416

Clarke, 2018, Body-shape diversity in Triassic–Early Cretaceous neopterygian fishes: sustained holostean disparity and predominantly gradual increases in teleost phenotypic variety, Paleobiology, 44, 402, 10.1017/pab.2018.8

Coates, 1999, Endocranial preservation of a Carboniferous actinopterygian from Lancashire, UK, and the interrelationships of primitive actinopterygians, Philosophical Transactions of the Royal Society of London. Series B: Biological Sciences, 354, 435, 10.1098/rstb.1999.0396

Cope, 1872, Observations on the systematic relations of the fishes, Proceedings of the American Association for the Advancement of Science, 20, 317

Cope, 1887, Zittel’s manual of palaeontology, American Naturalist, 21, 1014

Deesri, 2016, A new Ginglymodi (Actinopterygii, Holostei) from the Late Jurassic–Early Cretaceous of Thailand, with comments on the early diversification of Lepisosteiformes in Southeast Asia, Journal of Vertebrate Paleontology, 36, e1225747, 10.1080/02724634.2016.1225747

Deesri, 2014, A new species of the ginglymodian fish Isanichthys from the Late Jurassic Phu Kradung Formation, northeastern Thailand, Acta Palaeontologica Polonica, 59, 313, 10.4202/app.2012.0013

Ebert, 2018, Cerinichthys koelblae, gen. et sp. nov., from the Upper Jurassic of Cerin, France, and its phylogenetic setting, leading to a reassessment of the phylogenetic relationships of Halecomorphi (Actinopterygii), Journal of Vertebrate Paleontology, 38, e1420071, 10.1080/02724634.2017.1420071

Friedman, 2015, The early evolution of ray-finned fishes, Palaeontology, 58, 213, 10.1111/pala.12150

Gardiner, 1960, A revision of certain actinopterygian and coelacanth fishes, chiefly from the Lower Lias, Bulletin of the British Museum (Natural History), Geology, 4, 239, 10.5962/p.313860

Gardiner, 1996, Interrelationships of basal neopterygians, Interrelationships of Fishes, 117, 10.1016/B978-012670950-6/50007-2

Gardiner, 1989, Interrelationships of lower actinopterygian fishes, Zoological Journal of the Linnean Society, 97, 135, 10.1111/j.1096-3642.1989.tb00550.x

Giles, 2017, Early members of ‘living fossil’ lineage imply later origin of modern ray-finned fishes, Nature, 549, 265, 10.1038/nature23654

Gill, 1923, The Permian fishes of the genus Acentrophorus, Proceedings of the Zoological Society of London, 93, 19, 10.1111/j.1096-3642.1923.tb02170.x

Goodrich, 1930, Studies on the structure and development of vertebrates, 10.5962/bhl.title.82144

Grande, 2010, An empirical synthetic pattern study of gars (Lepisosteiformes) and closely related species, based mostly on skeletal anatomy. The resurrection of Holostei, Copeia, 10, 1

Grande, 1998, A comprehensive phylogenetic study of amiid fishes (Amiidae) based on comparative skeletal anatomy: an empirical search for interconnected patterns of natural history, Society of Vertebrate Paleontology Memoir (supplement to Journal of Vertebrate Paleontology), 18, 1, 10.1080/02724634.1998.10011114

Hurley, 2007, A new time-scale for ray-finned fish evolution, Proceedings of the Royal Society B: Biological Sciences, 274, 489, 10.1098/rspb.2006.3749

Huxley, 1861, Preliminary essay upon the systematic arrangement of the fishes of the Devonian epoch, Memoirs of the Geological Survey of the United Kingdom, 10, 1

Inoue, 2009, The historical biogeography of the freshwater knifefishes using mitogenomic approaches: a Mesozoic origin of the Asian notopterids (Actinopterygii: Osteoglossomorpha), Molecular Phylogenetics and Evolution, 51, 486, 10.1016/j.ympev.2009.01.020

Lambers, 1992, On the Ichthyofauna of the Solnhofen Lithographic Limestone (Upper Jurassic, Germany)

Lane, 2012, Revision of Furo muensteri (Halecomorphi, Ophiopsidae) from the Upper Jurassic of western Europe, with comments on the genus, Journal of Vertebrate Paleontology, 32, 799, 10.1080/02724634.2012.680325

Lane, 2015, A taxonomic reassessment of Ophiopsis (Halecomorphi, Ionoscopiformes), with a revision of Upper Jurassic species from the Solnhofen Archipelago, and a new genus of Ophiopsidae, Journal of Vertebrate Paleontology, 35, e883238, 10.1080/02724634.2014.883238

Liem, 2001, Functional anatomy of the vertebrates: an evolutionary perspective, 3

Liu, 2002, On the most primitive amiid fish from Upper Triassic of Xingyi, Guizhou, Acta Palaeontologica Sinica, 41, 461

Liu, 2003, New discovered fishes from Keichousaurus bearing horizon of Late Triassic in Xingyi of Guizhou, Acta Palaeontologica Sinica, 42, 346

Lombardo, 2001, Actinopterygians from the Middle Triassic of northern Italy and Canton Ticino (Switzerland): anatomical descriptions and nomenclatural problems, Rivista Italiana di Paleontologia e Stratigrafia, 107, 345, 10.13130/2039-4942/5439

López-Arbarello, 2012, Phylogenetic interrelationships of ginglymodian fishes (Actinopterygii: Neopterygii), PLOS ONE, 7, e39370, 10.1371/journal.pone.0039370

López-Arbarello, 2016, New holostean fishes (Actinopterygii: Neopterygii) from the Middle Triassic of the Monte San Giorgio (Canton Ticino, Switzerland), PeerJ, 4, e2234, 10.7717/peerj.2234

López-Arbarello, 2019, Taxonomy and phylogeny of Eosemionotus Stolley, 1920 (Neopterygii: Ginglymodi) from the Middle Triassic of Europe, Palaeontologia Electronica, 22.1.10A, 1, 10.26879/904

López-Arbarello, 2018, Neopterygian phylogeny: the merger assay, Royal Society Open Science, 5, 172337, 10.1098/rsos.172337

López-Arbarello, 2014, Phylogenetic relationships of the Triassic Archaeosemionotus Deecke (Halecomorphi, Ionoscopiformes) from the ‘Perledo Fauna’, PLOS ONE, 9, e108665, 10.1371/journal.pone.0108665

López-Arbarello, 2011, New species of Sangiorgioichthys Tintori and Lombardo, 2007 (Neopterygii, Semionotiformes) from the Anisian of Luoping (Yunnan Province, South China), Zootaxa, 2749, 25, 10.11646/zootaxa.2749.1.2

Ma, 2017, A new ionoscopiform fish (Holostei: Halecomorphi) from the Middle Triassic (Anisian) of Yunnan, China, Vertebrata PalAsiatica, 55, 92

Machado, 2013, Teoichthys brevipina, sp. nov., a new ophiopsid fish (Halecomorphi, Ionoscopiformes) from the Lower Cretaceous Tlayúa Formation, Central Mexico, Journal of Vertebrate Paleontology, 33, 482, 10.1080/02724634.2013.729962

Maisey, 1991, Oshunia Wenz and Kellner, 1986, Santana Fossils: An Illustrated Atlas, 157

Müller, 1844, Über den Bau und die Grenzen der Ganoiden und über das natürliche System der Fishe, Bericht Akademie der Wiessenschaften Berlin, 1844, 416

Nakatani, 2011, Evolutionary history of Otophysi (Teleostei), a major clade of the modern freshwater fishes: Pangaean origin and Mesozoic radiation, BMC Evolutionary Biology, 11, 177, 10.1186/1471-2148-11-177

Near, 2012, Resolution of ray-finned fish phylogeny and timing of diversification, Proceedings of the National Academy of Sciences of the United States of America, 109, 13698, 10.1073/pnas.1206625109

Nelson, 1969, Infraorbital bones and their bearing on the phylogeny and geography of osteoglossomorph fishes, American Museum Novitates, 2394, 1

Nielsen, 1942, Studies on Triassic fishes from East Greenland. I. Glaucolepis and Boreosomus, Meddelser øm Grønland, 138, 1

Nielsen, 1949, Studies on Triassic fishes from East Greenland. II. Australosomus and Birgeria, Meddelser øm Grønland, 146, 1

Nursall, 1991, Neoproscinetes, Santana Fossils: An Illustrated Atlas, 124

Olsen, 1984, The skull and pectoral girdle of the parasemionotid fish Watsonulus eugnathoides from the Early Triassic Sakamena Group of Madagascar, with comments on the relationship of the holostean fishes, Journal of Vertebrate Paleontology, 4, 481, 10.1080/02724634.1984.10012024

Olsen, 1991, Morphology of the Semionotus elegans species group from the Early Jurassic part of the Newark supergroup of eastern North America with comments on the family Semionotidae (Neopterygii), Journal of Vertebrate Paleontology, 11, 269, 10.1080/02724634.1991.10011398

Patterson, 1973, Interrelationships of holosteans, Interrelationships of Fishes, 233

Regan, 1923, The skeleton of Lepidosteus, with remarks on the origin and evolution of the lower neopterygian fishes, Proceedings of the Zoological Society of London, 93, 445, 10.1111/j.1096-3642.1923.tb02191.x

Romer, 1945, Vertebrate paleontology, 2

Rosen, 1981, Lungfishes, tetrapods, paleontology, and plesiomorphy, Bulletin of the American Museum of Natural History, 167, 159

Sallan, 2014, Major issues in the origins of ray-finned fish (Actinopterygii) biodiversity, Biological Reviews, 89, 950, 10.1111/brv.12086

Schultze, 1966, Morphologische und histologische Untersuchungen an Schuppen mesozoischer Actinopterygier (Übergang von Ganoid- zu Rundschuppen), Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 126, 232

Schultze, 2015, Scales, enamel, cosmine, ganoine, and early Osteichthyans, Comptes Rendus Palevol, 15, 83, 10.1016/j.crpv.2015.04.001

Setiamarga, 2009, Divergence time of the two regional medaka populations in Japan as a new time scale for comparative genomics of vertebrates, Biological Letters, 5, 812, 10.1098/rsbl.2009.0419

Su, 1959, Triassic fishes from Kueichow, Southwest China, Vertebrata PalAsiatica, 3, 205

Sun, 2018, Revision of Kyphosichthys grandei Xu & Wu, 2012 from the Middle Triassic of Yunnan Province, South China: implications for phylogenetic interrelationships of ginglymodian fishes, Journal of Systematic Palaeontology, 16, 67, 10.1080/14772019.2016.1269049

Sun, 2017, A new non-parasemionotiform order of the Halecomorphi (Neopterygii, Actinopterygii) from the Middle Triassic of Tethys, Journal of Systematic Palaeontology, 15, 223, 10.1080/14772019.2016.1181679

Swofford, 2003, PAUP*. Phylogenetic analysis using parsimony (*and other methods)

Taverne, 2015, Osteology and phylogenetic relationships of Congophiopsis lepersonnei gen. nov. (Halecomorphi, Ionoscopiformes) from the Songa Limestones (Middle Jurassic, Stanleyville Formation), Democratic Republic of Congo, Geo-Eco-Trop, 38, 223

Tintori, 2010, A new basal neopterygian from the Middle Triassic of Luoping County (South China), Rivista Italiana di Paleontologia e Stratigrafia, 116, 161, 10.13130/2039-4942/5947

Tintori, 2015, Oldest stem Teleostei from the late Ladinian (Middle Triassic) of southern China, Rivista Italiana di Paleontologia e Stratigrafia, 121, 285, 10.13130/2039-4942/6519

Wen, 2012, A new genus of basal actinopterygian fish from the Anisian (Middle Triassic) of Luoping, Yunnan Province, Southwest China, Acta Palaeontology Polonica, 57, 149, 10.4202/app.2010.0089

Westoll, 1944, The Haplolepidae, a new family of Late Carboniferous bony fishes—a study in taxonomy and evolution, Bulletin of the American Museum of Natural History, 83, 1

Xu, 2011, A new scanilepiform from the Lower Triassic of northern Gansu Province, China, and phylogenetic relationships of non-teleostean Actinopterygii, Zoological Journal of the Linnean Society, 161, 595, 10.1111/j.1096-3642.2010.00645.x

Xu, 2015, Taxonomic revision of Plesiofuro mingshuica from the Lower Triassic of northern Gansu, China, and the relationships of early neopterygian clades, Journal of Vertebrate Paleontology, 35, e1001515, 10.1080/02724634.2014.1001515

Xu, 2016, A Middle Triassic stem-neopterygian fish from China sheds new light on the peltopleuriform phylogeny and internal fertilization, Science Bulletin, 61, 1766, 10.1007/s11434-016-1189-5

Xu, 2018, Redescription and phylogenetic reassessment of Asialepidotus shingyiensis (Holostei: Halecomorphi) from the Middle Triassic (Ladinian) of China, Zoological Journal of the Linnean Society, 184, 95, 10.1093/zoolinnean/zlx105

Xu, 2018a, Fuyuanichthys wangi gen. et sp. nov. from the Middle Triassic (Ladinian) of China highlights the early diversification of ginglymodian fishes, PeerJ, 6, e6054, 10.7717/peerj.6054

Xu, 2019, A Middle Triassic kyphosichthyiform from Yunnan, China, and phylogenetic reassessment of early ginglymodians, Vertebrata PalAsiatica, 57, 1, 10.19615/j.cnki.1000-3118.190319

Xu, 2018b, A large peltopleurid fish from the Middle Triassic (Ladinian) of Yunnan and Guizhou, China, Vertebrata PalAsiatica, 56, 106

Xu, 2015, Panxianichthys imparilis gen. et sp. nov., a new ionoscopiform (Halecomorphi) from the Middle Triassic of Guizhou, China, Vertebrata PalAsiatica, 53, 1

Xu, 2012, A deep-bodied ginglymodian fish from the Middle Triassic of eastern Yunnan Province, China, and the phylogeny of lower neopterygians, Chinese Science Bulletin, 57, 111, 10.1007/s11434-011-4719-1

Xu, 2016, A Middle Triassic stem-neopterygian fish from China shows remarkable secondary sexual characteristics, Science Bulletin, 61, 338, 10.1007/s11434-016-1007-0

Xu, 2014, The oldest ionoscopiform from China sheds new light on the early evolution of halecomorph fishes, Biology Letters, 10, 20140204, 10.1098/rsbl.2014.0204

Xu, 2012, A new stem-neopterygian fish from the Middle Triassic of China shows the earliest over-water gliding strategy of the vertebrates, Proceedings of the Royal Society B: Biological Sciences, 280, 20122261, 10.1098/rspb.2012.2261

Zhang, 2009, A conodont-based Middle Triassic age assignment for the Luoping Biota of Yunnan, China, Science in China Series D: Earth Sciences, 52, 1673, 10.1007/s11430-009-0114-z