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We apologize to those whose work was not cited because of space constraints. We thank C. Brühl D. L. Buhl I. Creese J. Csicsvari W. J. Freeman K. D. Harris D. A. Henze N. Kopell N. Maier A. Sirota X-J. Wang and M. Zugaro for discussions. Supported by the National Institute of Mental Health and the National Institute of Neurological Disorders and Stroke.