1Department of Biology, New York University, 100 Washington Square East, New York, New York 10003, USA
2CNRS-Institut Jacques Monod, 15 rue Hélène Brion, 75205, Paris cedex 13, France
3Department of Biology University of Utah Salt Lake City, Utah 84112 USA
4Institute of Developmental Biology and Cancer, CNRS-University of Nice, Sophia-Antipolis, Parc Valrose, 06108, NICE cedex 2, France