Frequency dependent corticofugal excitation of principal cells in the cat's dorsal lateral geniculate nucleus

Springer Science and Business Media LLC - Tập 79 - Trang 313-318 - 1990
S. Lindström1, A. Wróbel1
1Department of Physiology, University of Göteborg, Göteborg, Sweden

Tóm tắt

Excitatory postsynaptic potentials (EPSPs) were evoked in principal cells of the cat's dorsal lateral geniculate nucleus by electrical stimulation of cortico-geniculate fibres. The EPSPs had a pronounced frequency sensitivity. They were barely detectable at stimulation frequencies below 3 Hz but increased dramatically in size at higher frequencies. At 30–50 Hz their amplitude typically exceeded that of EPSPs from optic tract fibres. A prominent EPSP potentiation was also obtained with pair pulse stimulation. The findings are discussed in relation to the hypothesis that the cortico-geniculate system serves as a variable gain regulator for the visual input to the cortex.

Tài liệu tham khảo

Ahlsén G (1984) Brain stem neurones with differential projection to functional subregions of the dorsal lateral geniculate complex in the cat. Neuroscience 12: 817–838 Ahlsén G, Lindström S (1982) Excitation of perigeniculate neurones via axon collaterals of principal cells. Brain Res 236: 447–481 Ahlsén G, Lindström S (1983) Corticofugal projection to perigeniculate neurones in the cat. Acta Physiol Scand 118: 181–184 Ahlsén G, Grant K. Lindström S (1982) Monosynaptic excitation of principal cells in the lateral geniculate nucleus by corticofugal fibres. Brain Res 234: 454–458 Ahlsén G, Lindström S, Lo F-S (1984) Inhibition from the brain stem of inhibitory interneurones of the cat's dorsal lateral geniculate nucleus. J Physiol (Lond) 347: 593–609 Ahlsén G, Lindström S, Lo F-S (1985) Interaction between inhibitory pathways to principal cells in the lateral geniculate nucleus of the cat. Exp Brain Res 58: 134–143 Crick F (1984) Function of the thalamic reticular complex: the searchlight hypothesis. Proc Natl Acad Sci USA 81: 4586–4590 Del Castillo J, Katz B (1954) Statistical factors involved in neuromuscular facilitation and depression. J Physiol (Lond) 124: 574–585 Dubin MW, Cleland BG (1977) Organization of visual inputs to interneurons of lateral geniculate nucleus of the cat. J Neurophysiol 40: 410–427 Ferster D, Lindström S (1983) An intracellular analysis of geniculo-cortical connectivity in area 17 of the cat. J Physiol (Lond) 342: 181–215 Ferster D, Lindström S (1985a) Augmenting responses evoked in area 17 of the cat by intracortical axon collaterals of corticogeniculate cells. J Physiol (Lond) 367: 217–232 Ferster D, Lindström S (1985b) Synaptic excitation of neurones in area 17 of the cat by intracortical axon collaterals of cortico-geniculate cells. J Physiol (Lond) 367: 233–252 Gilbert CD, Kelly JP (1975) The projection of cells in different layers of the cat's visual cortex. J Comp Neurol 163: 81–106 Koch C (1987) The action of the corticofugal pathway on sensory thalamic nuclei: a hypothesis. Neurosci 23: 399–406 Lindström S (1982) Synaptic organization of inhibitory pathways to principal cells in the lateral geniculate nucleus of the cat. Brain Res 234: 447–453 Lindström S (1983) Interneurones in the lateral geniculate nucleus with monosynaptic excitation from retinal ganglion cells. Acta Physiol Scand 119: 101–103 Lindström S, Wróbel A (1985) Frequency enhancement of corticofugal excitation in the lateral geniculate nucleus of the cat. Neurosci Lett [Suppl] 22: S311 Livingstone MS, Hubel DH (1981) Effects of sleep and arousal on the processing of visual information in the cat. Nature 291: 554–561 Lømo T (1971) Potentiation of monosynaptic EPSPs in the perforant path-dentate granule cell synapse. Exp Brain Res 12: 46–63 Magleby KL (1973) The effect of tetanic and post-tetanic potentiation on facilitation of transmitter release at the frog neuromuscular junction. J Physiol (Lond) 234: 353–371 McNaughton BL (1980) Evidence for two physiologically distinct perforant pathways to the fascia dentata. Brain Res 199: 1–19 Murphy PC, Sillito AM (1987) Corticofugal feedback influences the generation of length tuning in the visual pathway. Nature 329: 727–729 Robson JA (1983) The morphology of corticofugal axons to the dorsal lateral geniculate nucleus in the cat. J Comp Neurol 216: 89–103 Schmielau F, Singer W (1977) The role of visual cortex for binocular interactions in the cat lateral geniculate nucleus. Brain Res 120: 354–361 Singer W (1973) The effect of mesencephalic reticular stimulation on intracellular potentials of cat lateral geniculate neurones. Brain Res 61: 35–54 Singer W (1977) Control of thalamic transmission by corticofugal and ascending reticular pathways in the visual system. Physiol Rev 57: 386–420 Singer W, Tretter F, Cynader M (1976) The effect of reticular stimulation on spontaneous and evoked activity in the visual cortex. Brain Res 102: 71–90 Tsumoto T, Suda K (1980) Three groups of cortico-geniculate neurons and their distribution in binocular and monocular segments of cat striate cortex. J Comp Neurol 193: 223–236 Tsumoto T, Creutzfeldt OD, Legendy CR (1978) Functional organization of the corticofugal system from visual cortex to lateral geniculate nucleus in the cat (with an appendix on geniculo-cortical monosynaptic connections). Exp Brain Res 32: 345–364