1Kobe University Research Center for Inland Seas, Rokkodai, Nadaku, Kobe 657-8501, Japan
2Creative Research Initiative 'Sousei', Hokkaido University, Sapporo, 060-0810 Japan
3Aichi Fisheries Research Institute, Gamagori 443-0021, Japan
4Asahimachi, Gamagori 443-8601, Japan