1Department of Veterinary Clinical Sciences, College of Veterinary Medicine, Iowa State University, Ames, Iowa, USA
2USDA-ARS, Ames, Iowa, USA
3Department of Small Animal Clinical Sciences, Gastrointestinal Laboratory, College of Veterinary Medicine and Biomedical Sciences, Texas A&M University, College Station, TX, USA
4School of Biosciences and Veterinary Medicine, University of Camerino, Camerino,Macerata, Italy
5Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, USA
6Department of Genetics, Development & Cell Biology, College of Liberal Arts and Sciences, Iowa State University, Ames, Iowa, USA
7Colorado State University Veterinary Teaching Hospital, Fort Collins, CO, USA
8Veterinary Specialty Hospital – San Diego, CA, USA
9Department of Veterinary Pathology, College of Veterinary Medicine, Iowa State University, Ames, Iowa, USA