1Shanghai Center for Plant Stress Biology, CAS Center for Excellence in Molecular Plant Sciences, Chinese Academy of Sciences, Shanghai 200032, China
2Department of Plant & Environmental Sciences, University of Copenhagen, 1871 Frederiksberg C, Denmark
3Copenhagen Plant Science Center, University of Copenhagen, 1871 Frederiksberg C, Denmark
4Joint International Research Laboratory of Metabolic & Developmental Sciences, State Key Laboratory of Hybrid Rice, SJTU-University of Adelaide Joint Centre for Agriculture and Health, School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, Shanghai, China