Pericytes: Developmental, Physiological, and Pathological Perspectives, Problems, and Promises

Developmental Cell - Tập 21 Số 2 - Trang 193-215 - 2011
Annika Keller1,2, Guillem Genové1, Christer Betsholtz1
1Division of Vascular Biology, Department of Medical Biochemistry and Biophysics, Karolinska Institute, SE-171 77 Stockholm, Sweden
2Institute of Neuropathology, University Hospital Zürich, CH-8091 Zürich, Switzerland

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Abbott, 2006, Astrocyte-endothelial interactions at the blood-brain barrier, Nat. Rev. Neurosci., 7, 41, 10.1038/nrn1824

Abbott, 2010, Structure and function of the blood-brain barrier, Neurobiol. Dis., 37, 13, 10.1016/j.nbd.2009.07.030

Abramsson, 2002, Analysis of mural cell recruitment to tumor vessels, Circulation, 105, 112, 10.1161/hc0102.101437

Abramsson, 2003, Endothelial and nonendothelial sources of PDGF-B regulate pericyte recruitment and influence vascular pattern formation in tumors, J. Clin. Invest., 112, 1142, 10.1172/JCI200318549

Abramsson, 2007, Defective N-sulfation of heparan sulfate proteoglycans limits PDGF-BB binding and pericyte recruitment in vascular development, Genes Dev., 21, 316, 10.1101/gad.398207

Aguirre, 2004, NG2-expressing cells in the subventricular zone are type C-like cells and contribute to interneuron generation in the postnatal hippocampus, J. Cell Biol., 165, 575, 10.1083/jcb.200311141

Al Ahmad, 2009, Maintaining blood-brain barrier integrity: pericytes perform better than astrocytes during prolonged oxygen deprivation, J. Cell. Physiol., 218, 612, 10.1002/jcp.21638

Allende, 2002, Sphingosine-1-phosphate receptors and the development of the vascular system, Biochim. Biophys. Acta, 1582, 222, 10.1016/S1388-1981(02)00175-0

Allende, 2003, G-protein-coupled receptor S1P1 acts within endothelial cells to regulate vascular maturation, Blood, 102, 3665, 10.1182/blood-2003-02-0460

Anderson, 2011, Angiogenic sprouting into neural tissue requires Gpr124, an orphan G protein-coupled receptor, Proc. Natl. Acad. Sci. USA, 108, 2807, 10.1073/pnas.1019761108

Andrae, 2008, Role of platelet-derived growth factors in physiology and medicine, Genes Dev., 22, 1276, 10.1101/gad.1653708

Anjos-Afonso, 2011, Prospective identification and isolation of murine bone marrow derived multipotent mesenchymal progenitor cells, Best Pract. Res. Clin. Haematol., 24, 13, 10.1016/j.beha.2010.11.003

Armulik, 2005, Endothelial/pericyte interactions, Circ. Res., 97, 512, 10.1161/01.RES.0000182903.16652.d7

Armulik, 2010, Pericytes regulate the blood-brain barrier, Nature, 468, 557, 10.1038/nature09522

Asahina, 2011, Septum transversum-derived mesothelium gives rise to hepatic stellate cells and perivascular mesenchymal cells in developing mouse liver, Hepatology, 53, 983, 10.1002/hep.24119

Attwell, 2010, Glial and neuronal control of brain blood flow, Nature, 468, 232, 10.1038/nature09613

Baluk, 2005, Cellular abnormalities of blood vessels as targets in cancer, Curr. Opin. Genet. Dev., 15, 102, 10.1016/j.gde.2004.12.005

Barber, 2011, The significance of vascular and neural apoptosis to the pathology of diabetic retinopathy, Invest. Ophthalmol. Vis. Sci., 52, 1156, 10.1167/iovs.10-6293

Bauer, 1998, Inhibition of angiogenesis by thalidomide requires metabolic activation, which is species-dependent, Biochem. Pharmacol., 55, 1827, 10.1016/S0006-2952(98)00046-X

Belachew, 2003, Postnatal NG2 proteoglycan-expressing progenitor cells are intrinsically multipotent and generate functional neurons, J. Cell Biol., 161, 169, 10.1083/jcb.200210110

Bell, 2010, Pericytes control key neurovascular functions and neuronal phenotype in the adult brain and during brain aging, Neuron, 68, 409, 10.1016/j.neuron.2010.09.043

Benjamin, 1999, Selective ablation of immature blood vessels in established human tumors follows vascular endothelial growth factor withdrawal, J. Clin. Invest., 103, 159, 10.1172/JCI5028

Berg, 1997, The activin receptor-like kinase 1 gene: genomic structure and mutations in hereditary hemorrhagic telangiectasia type 2, Am. J. Hum. Genet., 61, 60, 10.1086/513903

Berger, 2005, Regulator of G-protein signaling-5 induction in pericytes coincides with active vessel remodeling during neovascularization, Blood, 105, 1094, 10.1182/blood-2004-06-2315

Bergers, 2003, Benefits of targeting both pericytes and endothelial cells in the tumor vasculature with kinase inhibitors, J. Clin. Invest., 111, 1287, 10.1172/JCI200317929

Bergwerff, 1998, Neural crest cell contribution to the developing circulatory system: implications for vascular morphology?, Circ. Res., 82, 221, 10.1161/01.RES.82.2.221

Bondjers, 2003, Transcription profiling of platelet-derived growth factor-B-deficient mouse embryos identifies RGS5 as a novel marker for pericytes and vascular smooth muscle cells, Am. J. Pathol., 162, 721, 10.1016/S0002-9440(10)63868-0

Bondjers, 2006, Microarray analysis of blood microvessels from PDGF-B and PDGF-Rbeta mutant mice identifies novel markers for brain pericytes, FASEB J., 20, 1703, 10.1096/fj.05-4944fje

Bresnick, 1977, Clinicopathologic correlations in diabetic retinopathy. II. Clinical and histologic appearances of retinal capillary microaneurysms, Arch. Ophthalmol., 95, 1215, 10.1001/archopht.1977.04450070113010

Cai, 2008, A myocardial lineage derives from Tbx18 epicardial cells, Nature, 454, 104, 10.1038/nature06969

Cai, 2008, The angiopoietin/Tie-2 system regulates pericyte survival and recruitment in diabetic retinopathy, Invest. Ophthalmol. Vis. Sci., 49, 2163, 10.1167/iovs.07-1206

Cambier, 2005, Integrin alpha(v)beta8-mediated activation of transforming growth factor-beta by perivascular astrocytes: an angiogenic control switch, Am. J. Pathol., 166, 1883, 10.1016/S0002-9440(10)62497-2

Canault, 2010, The lack of ADAM17 activity during embryonic development causes hemorrhage and impairs vessel formation, PLoS ONE, 5, e13433, 10.1371/journal.pone.0013433

Cardoso, 2010, Looking at the blood-brain barrier: molecular anatomy and possible investigation approaches, Brain Res. Brain Res. Rev., 64, 328, 10.1016/j.brainresrev.2010.05.003

Casanovas, 2011, The adaptive stroma joining the antiangiogenic resistance front, J. Clin. Invest., 121, 1244, 10.1172/JCI46430

Cascone, 2011, Upregulated stromal EGFR and vascular remodeling in mouse xenograft models of angiogenesis inhibitor-resistant human lung adenocarcinoma, J. Clin. Invest., 121, 1313, 10.1172/JCI42405

Cataldi, 2004, Imatinib-mesylate blocks recombinant T-type calcium channels expressed in human embryonic kidney-293 cells by a protein tyrosine kinase-independent mechanism, J. Pharmacol. Exp. Ther., 309, 208, 10.1124/jpet.103.061184

Chang, 1999, Smad5 knockout mice die at mid-gestation due to multiple embryonic and extraembryonic defects, Development, 126, 1631, 10.1242/dev.126.8.1631

Chappell, 2010, Vascular development: genetic mechanisms and links to vascular disease, Curr. Top. Dev. Biol., 90, 43, 10.1016/S0070-2153(10)90002-1

Chen, 2003, Smad proteins regulate transcriptional induction of the SM22alpha gene by TGF-beta, Nucleic Acids Res., 31, 1302, 10.1093/nar/gkg224

Chen, 2007, Spatio-temporal VEGF and PDGF delivery patterns blood vessel formation and maturation, Pharm. Res., 24, 258, 10.1007/s11095-006-9173-4

Cheung, 2010, Diabetic retinopathy, Lancet, 376, 124, 10.1016/S0140-6736(09)62124-3

Cho, 2003, Pericyte-specific expression of Rgs5: implications for PDGF and EDG receptor signaling during vascular maturation, FASEB J., 17, 440, 10.1096/fj.02-0340fje

Christian, 2008, Endosialin (Tem1) is a marker of tumor-associated myofibroblasts and tumor vessel-associated mural cells, Am. J. Pathol., 172, 486, 10.2353/ajpath.2008.070623

Clark, 1925, The development of adventitial (Rouget) cells on the blood capillaries of amphibian larvae, Am. J. Anat., 35, 239, 10.1002/aja.1000350205

Collett, 2003, Receptor tyrosine kinase Axl modulates the osteogenic differentiation of pericytes, Circ. Res., 92, 1123, 10.1161/01.RES.0000074881.56564.46

Corselli, 2010, Perivascular ancestors of adult multipotent stem cells, Arterioscler. Thromb. Vasc. Biol., 30, 1104, 10.1161/ATVBAHA.109.191643

Crisan, 2008, Purification and long-term culture of multipotent progenitor cells affiliated with the walls of human blood vessels: myoendothelial cells and pericytes, Methods Cell Biol., 86, 295, 10.1016/S0091-679X(08)00013-7

Crisan, 2008, A perivascular origin for mesenchymal stem cells in multiple human organs, Cell Stem Cell, 3, 301, 10.1016/j.stem.2008.07.003

Cullen, 2011, GPR124, an orphan G protein-coupled receptor, is required for CNS-specific vascularization and establishment of the blood-brain barrier, Proc. Natl. Acad. Sci. USA, 108, 5759, 10.1073/pnas.1017192108

Dai, 2009, Visualization and contractile activity of cochlear pericytes in the capillaries of the spiral ligament, Hear. Res., 254, 100, 10.1016/j.heares.2009.04.018

Daneman, 2010, Pericytes are required for blood-brain barrier integrity during embryogenesis, Nature, 468, 562, 10.1038/nature09513

Davidoff, 2004, Progenitor cells of the testosterone-producing Leydig cells revealed, J. Cell Biol., 167, 935, 10.1083/jcb.200409107

Davis, 1996, Isolation of angiopoietin-1, a ligand for the TIE2 receptor, by secretion-trap expression cloning, Cell, 87, 1161, 10.1016/S0092-8674(00)81812-7

De Bock, 2011, Vessel abnormalization: another hallmark of cancer? Molecular mechanisms and therapeutic implications, Curr. Opin. Genet. Dev., 21, 73, 10.1016/j.gde.2010.10.008

De Palma, 2005, Tie2 identifies a hematopoietic lineage of proangiogenic monocytes required for tumor vessel formation and a mesenchymal population of pericyte progenitors, Cancer Cell, 8, 211, 10.1016/j.ccr.2005.08.002

Dellavalle, 2007, Pericytes of human skeletal muscle are myogenic precursors distinct from satellite cells, Nat. Cell Biol., 9, 255, 10.1038/ncb1542

Dente, 2001, Pericytes augment the capillary barrier in in vitro cocultures, J. Surg. Res., 97, 85, 10.1006/jsre.2001.6117

Dermietzel, 1991, Molecular anatomy of the blood-brain barrier as defined by immunocytochemistry, Int. Rev. Cytol., 127, 57, 10.1016/S0074-7696(08)60692-0

Dettman, 1998, Common epicardial origin of coronary vascular smooth muscle, perivascular fibroblasts, and intermyocardial fibroblasts in the avian heart, Dev. Biol., 193, 169, 10.1006/dbio.1997.8801

Díaz-Flores, 2009, Pericytes. Morphofunction, interactions and pathology in a quiescent and activated mesenchymal cell niche, Histol. Histopathol., 24, 909

Dickson, 1995, Defective haematopoiesis and vasculogenesis in transforming growth factor-beta 1 knock out mice, Development, 121, 1845, 10.1242/dev.121.6.1845

Ding, 2010, Inductive angiocrine signals from sinusoidal endothelium are required for liver regeneration, Nature, 468, 310, 10.1038/nature09493

Doherty, 1998, Vascular pericytes express osteogenic potential in vitro and in vivo, J. Bone Miner. Res., 13, 828, 10.1359/jbmr.1998.13.5.828

Dohgu, 2005, Brain pericytes contribute to the induction and up-regulation of blood-brain barrier functions through transforming growth factor-beta production, Brain Res., 1038, 208, 10.1016/j.brainres.2005.01.027

Domenga, 2004, Notch3 is required for arterial identity and maturation of vascular smooth muscle cells, Genes Dev., 18, 2730, 10.1101/gad.308904

Dore-Duffy, 2006, CNS microvascular pericytes exhibit multipotential stem cell activity, J. Cereb. Blood Flow Metab., 26, 613, 10.1038/sj.jcbfm.9600272

Drake, 1998, Morphogenesis of the first blood vessels, Ann. N Y Acad. Sci., 857, 155, 10.1111/j.1749-6632.1998.tb10115.x

Du, 2008, HIF1alpha induces the recruitment of bone marrow-derived vascular modulatory cells to regulate tumor angiogenesis and invasion, Cancer Cell, 13, 206, 10.1016/j.ccr.2008.01.034

Duffield, 2011, Origin of new cells in the adult kidney: results from genetic labeling techniques, Kidney Int., 79, 494, 10.1038/ki.2010.338

Dumont, 1993, The endothelial-specific receptor tyrosine kinase, tek, is a member of a new subfamily of receptors, Oncogene, 8, 1293

Dumont, 1994, Dominant-negative and targeted null mutations in the endothelial receptor tyrosine kinase, tek, reveal a critical role in vasculogenesis of the embryo, Genes Dev., 8, 1897, 10.1101/gad.8.16.1897

Eberth, C.J. (1871). Handbuch der Lehre von der Gewegen des Menschen und der Tiere., Vol 1 (Leipzig).

Ehninger, 2011, The bone marrow stem cell niche grows up: mesenchymal stem cells and macrophages move in, J. Exp. Med., 208, 421, 10.1084/jem.20110132

Ehrlich, 2010, Diabetic macular oedema: physical, physiological and molecular factors contribute to this pathological process, Acta Ophthalmol. (Copenh.), 88, 279, 10.1111/j.1755-3768.2008.01501.x

Enge, 2002, Endothelium-specific platelet-derived growth factor-B ablation mimics diabetic retinopathy, EMBO J., 21, 4307, 10.1093/emboj/cdf418

Enzinger, 1976, Hemangiopericytoma. An analysis of 106 cases, Hum. Pathol., 7, 61, 10.1016/S0046-8177(76)80006-8

Erber, 2004, Combined inhibition of VEGF and PDGF signaling enforces tumor vessel regression by interfering with pericyte-mediated endothelial cell survival mechanisms, FASEB J., 18, 338, 10.1096/fj.03-0271fje

Ergün, 2011, Vascular wall as a reservoir for different types of stem and progenitor cells, Antioxid. Redox Signal., 15, 981, 10.1089/ars.2010.3507

Etchevers, 2001, The cephalic neural crest provides pericytes and smooth muscle cells to all blood vessels of the face and forebrain, Development, 128, 1059, 10.1242/dev.128.7.1059

Fabian, 2005, A small molecule-kinase interaction map for clinical kinase inhibitors, Nat. Biotechnol., 23, 329, 10.1038/nbt1068

Fabris, 2011, Epithelial-mesenchymal interactions in biliary diseases, Semin. Liver Dis., 31, 11, 10.1055/s-0031-1272832

Falcón, 2009, Contrasting actions of selective inhibitors of angiopoietin-1 and angiopoietin-2 on the normalization of tumor blood vessels, Am. J. Pathol., 175, 2159, 10.2353/ajpath.2009.090391

Farrington-Rock, 2004, Chondrogenic and adipogenic potential of microvascular pericytes, Circulation, 110, 2226, 10.1161/01.CIR.0000144457.55518.E5

Feng, 2011, Dual origin of mesenchymal stem cells contributing to organ growth and repair, Proc. Natl. Acad. Sci. USA, 108, 6503, 10.1073/pnas.1015449108

Fernández-Klett, 2010, Pericytes in capillaries are contractile in vivo, but arterioles mediate functional hyperemia in the mouse brain, Proc. Natl. Acad. Sci. USA, 107, 22290, 10.1073/pnas.1011321108

Fletcher, 2006, The evolving classification of soft tissue tumours: an update based on the new WHO classification, Histopathology, 48, 3, 10.1111/j.1365-2559.2005.02284.x

Foo, 2006, Ephrin-B2 controls cell motility and adhesion during blood-vessel-wall assembly, Cell, 124, 161, 10.1016/j.cell.2005.10.034

Foster, 2008, Contribution of neural crest-derived cells in the embryonic and adult thymus, J. Immunol., 180, 3183, 10.4049/jimmunol.180.5.3183

Friedenstein, 1970, The development of fibroblast colonies in monolayer cultures of guinea-pig bone marrow and spleen cells, Cell Tissue Kinet., 3, 393

Furuhashi, 2004, Platelet-derived growth factor production by B16 melanoma cells leads to increased pericyte abundance in tumors and an associated increase in tumor growth rate, Cancer Res., 64, 2725, 10.1158/0008-5472.CAN-03-1489

Gaengel, 2009, Endothelial-mural cell signaling in vascular development and angiogenesis, Arterioscler. Thromb. Vasc. Biol., 29, 630, 10.1161/ATVBAHA.107.161521

Gallione, 2004, A combined syndrome of juvenile polyposis and hereditary haemorrhagic telangiectasia associated with mutations in MADH4 (SMAD4), Lancet, 363, 852, 10.1016/S0140-6736(04)15732-2

Galvez, 2008, Cardiac mesoangioblasts are committed, self-renewable progenitors, associated with small vessels of juvenile mouse ventricle, Cell Death Differ., 15, 1417, 10.1038/cdd.2008.75

García-Gómez, 2010, Mesenchymal stem cells: biological properties and clinical applications, Expert Opin. Biol. Ther., 10, 1453, 10.1517/14712598.2010.519333

Gardner, 2011, An integrated approach to diabetic retinopathy research, Arch. Ophthalmol., 129, 230, 10.1001/archophthalmol.2010.362

Gengler, 2006, Solitary fibrous tumour and haemangiopericytoma: evolution of a concept, Histopathology, 48, 63, 10.1111/j.1365-2559.2005.02290.x

Geraldes, 2009, Activation of PKC-delta and SHP-1 by hyperglycemia causes vascular cell apoptosis and diabetic retinopathy, Nat. Med., 15, 1298, 10.1038/nm.2052

Gerhardt, 2003, Endothelial-pericyte interactions in angiogenesis, Cell Tissue Res., 314, 15, 10.1007/s00441-003-0745-x

Gerhardt, 2008, Pericytes: gatekeepers in tumour cell metastasis?, J. Mol. Med., 86, 135, 10.1007/s00109-007-0258-2

Gerhardt, 2000, N-cadherin mediates pericytic-endothelial interaction during brain angiogenesis in the chicken, Dev. Dyn., 218, 472, 10.1002/1097-0177(200007)218:3<472::AID-DVDY1008>3.0.CO;2-#

Gnessi, 2000, Leydig cell loss and spermatogenic arrest in platelet-derived growth factor (PDGF)-A-deficient mice, J. Cell Biol., 149, 1019, 10.1083/jcb.149.5.1019

Goossens, 2011, The EMT regulator Zeb2/Sip1 is essential for murine embryonic hematopoietic stem/progenitor cell differentiation and mobilization, Blood, 117, 5620, 10.1182/blood-2010-08-300236

Goumans, 2002, Balancing the activation state of the endothelium via two distinct TGF-beta type I receptors, EMBO J., 21, 1743, 10.1093/emboj/21.7.1743

Goumans, 2003, Activin receptor-like kinase (ALK)1 is an antagonistic mediator of lateral TGFbeta/ALK5 signaling, Mol. Cell, 12, 817, 10.1016/S1097-2765(03)00386-1

Greenberg, 2008, A role for VEGF as a negative regulator of pericyte function and vessel maturation, Nature, 456, 809, 10.1038/nature07424

Hainsworth, 2005, Treatment of metastatic renal cell carcinoma with a combination of bevacizumab and erlotinib, J. Clin. Oncol., 23, 7889, 10.1200/JCO.2005.01.8234

Hainsworth, 2007, Treatment of advanced renal cell carcinoma with the combination bevacizumab/erlotinib/imatinib: a phase I/II trial, Clin. Genitourin. Cancer, 5, 427, 10.3816/CGC.2007.n.030

Hamilton, 2010, Pericyte-mediated regulation of capillary diameter: a component of neurovascular coupling in health and disease, Front Neuroenergetics, 2, 5, 10.3389/fnene.2010.00005

Hammes, 2002, Pericytes and the pathogenesis of diabetic retinopathy, Diabetes, 51, 3107, 10.2337/diabetes.51.10.3107

Hammes, 2004, Angiopoietin-2 causes pericyte dropout in the normal retina: evidence for involvement in diabetic retinopathy, Diabetes, 53, 1104, 10.2337/diabetes.53.4.1104

Hamzah, 2008, Vascular normalization in Rgs5-deficient tumours promotes immune destruction, Nature, 453, 410, 10.1038/nature06868

Hanahan, 2011, Hallmarks of cancer: the next generation, Cell, 144, 646, 10.1016/j.cell.2011.02.013

Hayashi, 2004, Effects of hypoxia on endothelial/pericytic co-culture model of the blood-brain barrier, Regul. Pept., 123, 77, 10.1016/j.regpep.2004.05.023

He, 2010, Gamma-secretase activating protein is a therapeutic target for Alzheimer's disease, Nature, 467, 95, 10.1038/nature09325

Heglind, 2005, Lack of the central nervous system- and neural crest-expressed forkhead gene Foxs1 affects motor function and body weight, Mol. Cell. Biol., 25, 5616, 10.1128/MCB.25.13.5616-5625.2005

Heldin, 1999, Mechanism of action and in vivo role of platelet-derived growth factor, Physiol. Rev., 79, 1283, 10.1152/physrev.1999.79.4.1283

Hellström, 1999, Role of PDGF-B and PDGFR-beta in recruitment of vascular smooth muscle cells and pericytes during embryonic blood vessel formation in the mouse, Development, 126, 3047, 10.1242/dev.126.14.3047

Hinz, 2007, Formation and function of the myofibroblast during tissue repair, J. Invest. Dermatol., 127, 526, 10.1038/sj.jid.5700613

Hirschi, 2003, Gap junction communication mediates transforming growth factor-beta activation and endothelial-induced mural cell differentiation, Circ. Res., 93, 429, 10.1161/01.RES.0000091259.84556.D5

Hoglund, 2010, Neointima formation: a local affair, Arterioscler. Thromb. Vasc. Biol., 30, 1877, 10.1161/ATVBAHA.110.211433

Hori, 2004, A pericyte-derived angiopoietin-1 multimeric complex induces occludin gene expression in brain capillary endothelial cells through Tie-2 activation in vitro, J. Neurochem., 89, 503, 10.1111/j.1471-4159.2004.02343.x

Huang, 2010, Pericyte deficiencies lead to aberrant tumor vascularizaton in the brain of the NG2 null mouse, Dev. Biol., 344, 1035, 10.1016/j.ydbio.2010.06.023

Hungerford, 1999, Developmental biology of the vascular smooth muscle cell: building a multilayered vessel wall, J. Vasc. Res., 36, 2, 10.1159/000025622

Iivanainen, 2003, Angiopoietin-regulated recruitment of vascular smooth muscle cells by endothelial-derived heparin binding EGF-like growth factor, FASEB J., 17, 1609, 10.1096/fj.02-0939com

Iwamoto, 2003, Heparin-binding EGF-like growth factor and ErbB signaling is essential for heart function, Proc. Natl. Acad. Sci. USA, 100, 3221, 10.1073/pnas.0537588100

Jain, 2005, Normalization of tumor vasculature: an emerging concept in antiangiogenic therapy, Science, 307, 58, 10.1126/science.1104819

Jeansson, 2011, Angiopoietin-1 is essential in mouse vasculature during development and in response to injury, J. Clin. Invest., 121, 2278, 10.1172/JCI46322

Jin, 2008, Notch signaling regulates platelet-derived growth factor receptor-beta expression in vascular smooth muscle cells, Circ. Res., 102, 1483, 10.1161/CIRCRESAHA.107.167965

Jones, 2001, Rescue of the early vascular defects in Tek/Tie2 null mice reveals an essential survival function, EMBO Rep., 2, 438, 10.1093/embo-reports/kve093

Joutel, 1996, Notch3 mutations in CADASIL, a hereditary adult-onset condition causing stroke and dementia, Nature, 383, 707, 10.1038/383707a0

Joyce, 2009, Microenvironmental regulation of metastasis, Nat. Rev. Cancer, 9, 239, 10.1038/nrc2618

Kern, 2000, Response of capillary cell death to aminoguanidine predicts the development of retinopathy: comparison of diabetes and galactosemia, Invest. Ophthalmol. Vis. Sci., 41, 3972

Kienast, 2010, Real-time imaging reveals the single steps of brain metastasis formation, Nat. Med., 16, 116, 10.1038/nm.2072

Kisseleva, 2008, Mechanisms of fibrogenesis, Exp. Biol. Med. (Maywood), 233, 109, 10.3181/0707-MR-190

Klaassen, 2009, Altered expression of genes related to blood-retina barrier disruption in streptozotocin-induced diabetes, Exp. Eye Res., 89, 4, 10.1016/j.exer.2009.01.006

Koch, 2008, Malignant tumors of blood vessels: angiosarcomas, hemangioendotheliomas, and hemangioperictyomas, J. Surg. Oncol., 97, 321, 10.1002/jso.20973

Korn, 2002, Neuroectodermal origin of brain pericytes and vascular smooth muscle cells, J. Comp. Neurol., 442, 78, 10.1002/cne.1423

Krueger, 2010, CNS pericytes: concepts, misconceptions, and a way out, Glia, 58, 1, 10.1002/glia.20898

Krüger, 2000, Defective vascular development in connexin 45-deficient mice, Development, 127, 4179, 10.1242/dev.127.19.4179

Kuhnert, 2010, Essential regulation of CNS angiogenesis by the orphan G protein-coupled receptor GPR124, Science, 330, 985, 10.1126/science.1196554

Kunz, 1994, The 140-kDa protein of blood-brain barrier-associated pericytes is identical to aminopeptidase N, J. Neurochem., 62, 2375, 10.1046/j.1471-4159.1994.62062375.x

Kurup, 2006, Heparan sulphate requirement in platelet-derived growth factor B-mediated pericyte recruitment, Biochem. Soc. Trans., 34, 454, 10.1042/BST0340454

Lai, 2005, The critical component to establish in vitro BBB model: Pericyte, Brain Res. Brain Res. Rev., 50, 258, 10.1016/j.brainresrev.2005.07.004

Lammert, 2001, Induction of pancreatic differentiation by signals from blood vessels, Science, 294, 564, 10.1126/science.1064344

Lamont, 2010, Hedgehog signaling via angiopoietin1 is required for developmental vascular stability, Mech. Dev., 127, 159, 10.1016/j.mod.2010.02.001

Lan, 2007, Essential role of endothelial Smad4 in vascular remodeling and integrity, Mol. Cell. Biol., 27, 7683, 10.1128/MCB.00577-07

Larson, 1987, Junctional transfer of small molecules in cultured bovine brain microvascular endothelial cells and pericytes, Microvasc. Res., 34, 184, 10.1016/0026-2862(87)90052-5

Larsson, 2001, Abnormal angiogenesis but intact hematopoietic potential in TGF-beta type I receptor-deficient mice, EMBO J., 20, 1663, 10.1093/emboj/20.7.1663

Lebrin, 2010, Thalidomide stimulates vessel maturation and reduces epistaxis in individuals with hereditary hemorrhagic telangiectasia, Nat. Med., 16, 420, 10.1038/nm.2131

Lee, 1999, Vascular endothelial cell adherens junction assembly and morphogenesis induced by sphingosine-1-phosphate, Cell, 99, 301, 10.1016/S0092-8674(00)81661-X

Lendahl, 1990, CNS stem cells express a new class of intermediate filament protein, Cell, 60, 585, 10.1016/0092-8674(90)90662-X

Levéen, 1994, Mice deficient for PDGF B show renal, cardiovascular, and hematological abnormalities, Genes Dev., 8, 1875, 10.1101/gad.8.16.1875

Li, 1999, Defective angiogenesis in mice lacking endoglin, Science, 284, 1534, 10.1126/science.284.5419.1534

Li, 2011, Endothelial Smad4 maintains cerebrovascular integrity by activating N-cadherin through cooperation with Notch, Dev. Cell, 20, 291, 10.1016/j.devcel.2011.01.011

Lin, 2008, Pericytes and perivascular fibroblasts are the primary source of collagen-producing cells in obstructive fibrosis of the kidney, Am. J. Pathol., 173, 1617, 10.2353/ajpath.2008.080433

Lindahl, 1997, Pericyte loss and microaneurysm formation in PDGF-B-deficient mice, Science, 277, 242, 10.1126/science.277.5323.242

Lindahl, 1998, Paracrine PDGF-B/PDGF-Rbeta signaling controls mesangial cell development in kidney glomeruli, Development, 125, 3313, 10.1242/dev.125.17.3313

Lindblom, 2003, Endothelial PDGF-B retention is required for proper investment of pericytes in the microvessel wall, Genes Dev., 17, 1835, 10.1101/gad.266803

Liu, 2000, Edg-1, the G protein-coupled receptor for sphingosine-1-phosphate, is essential for vascular maturation, J. Clin. Invest., 106, 951, 10.1172/JCI10905

Liu, 2009, NOTCH3 expression is induced in mural cells through an autoregulatory loop that requires endothelial-expressed JAGGED1, Circ. Res., 104, 466, 10.1161/CIRCRESAHA.108.184846

Liu, 2010, Notch3 is critical for proper angiogenesis and mural cell investment, Circ. Res., 107, 860, 10.1161/CIRCRESAHA.110.218271

Louis, 2007, The 2007 WHO classification of tumours of the central nervous system, Acta Neuropathol., 114, 97, 10.1007/s00401-007-0243-4

Maes, 2010, Osteoblast precursors, but not mature osteoblasts, move into developing and fractured bones along with invading blood vessels, Dev. Cell, 19, 329, 10.1016/j.devcel.2010.07.010

Mahoney, 2011, A unifying hypothesis for scleroderma: identifying a target cell for scleroderma, Curr. Rheumatol. Rep., 13, 28, 10.1007/s11926-010-0152-8

Majesky, 2007, Developmental basis of vascular smooth muscle diversity, Arterioscler. Thromb. Vasc. Biol., 27, 1248, 10.1161/ATVBAHA.107.141069

Majesky, 2011, Vascular smooth muscle progenitor cells: building and repairing blood vessels, Circ. Res., 108, 365, 10.1161/CIRCRESAHA.110.223800

Mathiisen, 2010, The perivascular astroglial sheath provides a complete covering of the brain microvessels: an electron microscopic 3D reconstruction, Glia, 58, 1094, 10.1002/glia.20990

Matsumoto, 2001, Liver organogenesis promoted by endothelial cells prior to vascular function, Science, 294, 559, 10.1126/science.1063889

McAllister, 1994, Endoglin, a TGF-beta binding protein of endothelial cells, is the gene for hereditary haemorrhagic telangiectasia type 1, Nat. Genet., 8, 345, 10.1038/ng1294-345

Mehta, 2006, Signaling mechanisms regulating endothelial permeability, Physiol. Rev., 86, 279, 10.1152/physrev.00012.2005

Mendelson, 2010, Stimulation of platelet-derived growth factor receptor beta (PDGFRbeta) activates ADAM17 and promotes metalloproteinase-dependent cross-talk between the PDGFRbeta and epidermal growth factor receptor (EGFR) signaling pathways, J. Biol. Chem., 285, 25024, 10.1074/jbc.M110.102566

Mikawa, 1996, Pericardial mesoderm generates a population of coronary smooth muscle cells migrating into the heart along with ingrowth of the epicardial organ, Dev. Biol., 174, 221, 10.1006/dbio.1996.0068

Montanez, 2009, Alpha-parvin controls vascular mural cell recruitment to vessel wall by regulating RhoA/ROCK signalling, EMBO J., 28, 3132, 10.1038/emboj.2009.295

Morikawa, 2002, Abnormalities in pericytes on blood vessels and endothelial sprouts in tumors, Am. J. Pathol., 160, 985, 10.1016/S0002-9440(10)64920-6

Mulamalla, 2008, Rare case of hemangiopericytoma responds to sunitinib, Transl. Res., 151, 129, 10.1016/j.trsl.2007.12.007

Müller, 2008, Neural crest origin of perivascular mesenchyme in the adult thymus, J. Immunol., 180, 5344, 10.4049/jimmunol.180.8.5344

Nag, 2011, Review: molecular pathogenesis of blood-brain barrier breakdown in acute brain injury, Neuropathol. Appl. Neurobiol., 37, 3, 10.1111/j.1365-2990.2010.01138.x

Nakagawa, 2009, A new blood-brain barrier model using primary rat brain endothelial cells, pericytes and astrocytes, Neurochem. Int., 54, 253, 10.1016/j.neuint.2008.12.002

Nanba, 2006, Loss of HB-EGF in smooth muscle or endothelial cell lineages causes heart malformation, Biochem. Biophys. Res. Commun., 350, 315, 10.1016/j.bbrc.2006.09.060

Nehls, 1993, The versatility of microvascular pericytes: from mesenchyme to smooth muscle?, Histochemistry, 99, 1, 10.1007/BF00268014

Nehls, 1992, Pericyte involvement in capillary sprouting during angiogenesis in situ, Cell Tissue Res., 270, 469, 10.1007/BF00645048

Netzer, 2003, Gleevec inhibits beta-amyloid production but not Notch cleavage, Proc. Natl. Acad. Sci. USA, 100, 12444, 10.1073/pnas.1534745100

Nielsen, 2010, Sonic hedgehog is required for vascular outgrowth in the hindbrain choroid plexus, Dev. Biol., 340, 430, 10.1016/j.ydbio.2010.01.032

Nisancioglu, 2008, Generation and characterization of rgs5 mutant mice, Mol. Cell. Biol., 28, 2324, 10.1128/MCB.01252-07

Nisancioglu, 2010, The absence of pericytes does not increase the sensitivity of tumor vasculature to vascular endothelial growth factor-A blockade, Cancer Res., 70, 5109, 10.1158/0008-5472.CAN-09-4245

Nolan-Stevaux, 2010, Differential contribution to neuroendocrine tumorigenesis of parallel Egfr signaling in cancer cells and pericytes, Genes Cancer, 1, 125, 10.1177/1947601909358722

Nombela-Arrieta, 2011, The elusive nature and function of mesenchymal stem cells, Nat. Rev. Mol. Cell Biol., 12, 126, 10.1038/nrm3049

Nourshargh, 2010, Breaching multiple barriers: leukocyte motility through venular walls and the interstitium, Nat. Rev. Mol. Cell Biol., 11, 366, 10.1038/nrm2889

Ockrim, 2010, Managing diabetic retinopathy, BMJ, 341, c5400, 10.1136/bmj.c5400

Oh, 2000, Activin receptor-like kinase 1 modulates transforming growth factor-beta 1 signaling in the regulation of angiogenesis, Proc. Natl. Acad. Sci. USA, 97, 2626, 10.1073/pnas.97.6.2626

Olson, 2011, PDGFRβ signaling regulates mural cell plasticity and inhibits fat development, Dev. Cell, 20, 815, 10.1016/j.devcel.2011.04.019

Oshima, 1996, TGF-beta receptor type II deficiency results in defects of yolk sac hematopoiesis and vasculogenesis, Dev. Biol., 179, 297, 10.1006/dbio.1996.0259

Ostman, 1991, Identification of a cell retention signal in the B-chain of platelet-derived growth factor and in the long splice version of the A-chain, Cell Regul., 2, 503, 10.1091/mbc.2.7.503

Ota, 2002, Targets of transcriptional regulation by two distinct type I receptors for transforming growth factor-beta in human umbilical vein endothelial cells, J. Cell. Physiol., 193, 299, 10.1002/jcp.10170

Ozawa, 2005, Angiogenesis with pericyte abnormalities in a transgenic model of prostate carcinoma, Cancer, 104, 2104, 10.1002/cncr.21436

Ozerdem, 2001, NG2 proteoglycan is expressed exclusively by mural cells during vascular morphogenesis, Dev. Dyn., 222, 218, 10.1002/dvdy.1200

Paik, 2004, Sphingosine 1-phosphate receptor regulation of N-cadherin mediates vascular stabilization, Genes Dev., 18, 2392, 10.1101/gad.1227804

Paquet-Fifield, 2009, A role for pericytes as microenvironmental regulators of human skin tissue regeneration, J. Clin. Invest., 119, 2795

Park, 2009, New insights into the hemangiopericytoma/solitary fibrous tumor spectrum of tumors, Curr. Opin. Oncol., 21, 327, 10.1097/CCO.0b013e32832c9532

Park, 2010, Inhibiting the VEGF-VEGFR pathway in angiosarcoma, epithelioid hemangioendothelioma, and hemangiopericytoma/solitary fibrous tumor, Curr. Opin. Oncol., 22, 351, 10.1097/CCO.0b013e32833aaad4

Park, 2011, Activity of temozolomide and bevacizumab in the treatment of locally advanced, recurrent, and metastatic hemangiopericytoma and malignant solitary fibrous tumor, Cancer, 10.1002/cncr.26098

Passino, 2007, Regulation of hepatic stellate cell differentiation by the neurotrophin receptor p75NTR, Science, 315, 1853, 10.1126/science.1137603

Passman, 2008, A sonic hedgehog signaling domain in the arterial adventitia supports resident Sca1+ smooth muscle progenitor cells, Proc. Natl. Acad. Sci. USA, 105, 9349, 10.1073/pnas.0711382105

Patan, 1998, TIE1 and TIE2 receptor tyrosine kinases inversely regulate embryonic angiogenesis by the mechanism of intussusceptive microvascular growth, Microvasc. Res., 56, 1, 10.1006/mvre.1998.2081

Peppiatt, 2006, Bidirectional control of CNS capillary diameter by pericytes, Nature, 443, 700, 10.1038/nature05193

Petrova, 2004, Defective valves and abnormal mural cell recruitment underlie lymphatic vascular failure in lymphedema distichiasis, Nat. Med., 10, 974, 10.1038/nm1094

Pietras, 2010, Hallmarks of cancer: interactions with the tumor stroma, Exp. Cell Res., 316, 1324, 10.1016/j.yexcr.2010.02.045

Polyak, 2009, Co-evolution of tumor cells and their microenvironment, Trends Genet., 25, 30, 10.1016/j.tig.2008.10.012

Powell, 2011, Mesenchymal cells of the intestinal lamina propria, Annu. Rev. Physiol., 73, 213, 10.1146/annurev.physiol.70.113006.100646

Puro, 2007, Physiology and pathobiology of the pericyte-containing retinal microvasculature: new developments, Microcirculation, 14, 1, 10.1080/10739680601072099

Qin, 2010, COUP-TFII regulates tumor growth and metastasis by modulating tumor angiogenesis, Proc. Natl. Acad. Sci. USA, 107, 3687, 10.1073/pnas.0914619107

Que, 2008, Mesothelium contributes to vascular smooth muscle and mesenchyme during lung development, Proc. Natl. Acad. Sci. USA, 105, 16626, 10.1073/pnas.0808649105

Rajantie, 2004, Adult bone marrow-derived cells recruited during angiogenesis comprise precursors for periendothelial vascular mural cells, Blood, 104, 2084, 10.1182/blood-2004-01-0336

Ramaswamy, 2003, A molecular signature of metastasis in primary solid tumors, Nat. Genet., 33, 49, 10.1038/ng1060

Raza, 2010, Pericytes and vessel maturation during tumor angiogenesis and metastasis, Am. J. Hematol., 85, 593, 10.1002/ajh.21745

Romeo, 2002, Activation of nuclear factor-kappaB induced by diabetes and high glucose regulates a proapoptotic program in retinal pericytes, Diabetes, 51, 2241, 10.2337/diabetes.51.7.2241

Rouget, 1873, Memoire sur le developpement, la structures et les proprietes des capillaires sanguins et lymphatiques, Archs Physiol Norm Pathol, 5, 603

Ruchoux, 1995, Systemic vascular smooth muscle cell impairment in cerebral autosomal dominant arteriopathy with subcortical infarcts and leukoencephalopathy, Acta Neuropathol., 89, 500, 10.1007/BF00571504

Ruiter, 1993, Angiogenesis in wound healing and tumor metastasis, Behring Inst. Mitt., Aug., 258

Sahin, 2004, Distinct roles for ADAM10 and ADAM17 in ectodomain shedding of six EGFR ligands, J. Cell Biol., 164, 769, 10.1083/jcb.200307137

Sainson, 2008, Regulation of angiogenesis by homotypic and heterotypic notch signalling in endothelial cells and pericytes: from basic research to potential therapies, Angiogenesis, 11, 41, 10.1007/s10456-008-9098-0

Salvucci, 2009, EphrinB reverse signaling contributes to endothelial and mural cell assembly into vascular structures, Blood, 114, 1707, 10.1182/blood-2008-12-192294

Santoro, 2009, Characterization of vascular mural cells during zebrafish development, Mech. Dev., 126, 638, 10.1016/j.mod.2009.06.1080

Sato, 1989, Inhibition of endothelial cell movement by pericytes and smooth muscle cells: activation of a latent transforming growth factor-beta 1-like molecule by plasmin during co-culture, J. Cell Biol., 109, 309, 10.1083/jcb.109.1.309

Scholten, 2010, Genetic labeling does not detect epithelial-to-mesenchymal transition of cholangiocytes in liver fibrosis in mice, Gastroenterology, 139, 987, 10.1053/j.gastro.2010.05.005

Schrimpf, 2011, Mechanisms of fibrosis: the role of the pericyte, Curr. Opin. Nephrol. Hypertens., 20, 297, 10.1097/MNH.0b013e328344c3d4

Sennino, 2007, Sequential loss of tumor vessel pericytes and endothelial cells after inhibition of platelet-derived growth factor B by selective aptamer AX102, Cancer Res., 67, 7358, 10.1158/0008-5472.CAN-07-0293

Shepro, 1993, Pericyte physiology, FASEB J., 7, 1031, 10.1096/fasebj.7.11.8370472

Shi, 2003, Perivascular niche of postnatal mesenchymal stem cells in human bone marrow and dental pulp, J. Bone Miner. Res., 18, 696, 10.1359/jbmr.2003.18.4.696

Shi, 2011, Latent TGF-β structure and activation, Nature, 474, 343, 10.1038/nature10152

Shojaei, 2008, Role of the microenvironment in tumor growth and in refractoriness/resistance to anti-angiogenic therapies, Drug Resist. Updat., 11, 219, 10.1016/j.drup.2008.09.001

Sims, 1986, The pericyte—a review, Tissue Cell, 18, 153, 10.1016/0040-8166(86)90026-1

Sims, 1994, Heterogeneity of pericyte populations in equine skeletal muscle and dermal microvessels: a quantitative study, Anat. Histol. Embryol., 23, 232, 10.1111/j.1439-0264.1994.tb00472.x

Smith, 2006, Endocytosis at the blood-brain barrier: from basic understanding to drug delivery strategies, J. Drug Target., 14, 191, 10.1080/10611860600650086

Soehnlein, 2010, Phagocyte partnership during the onset and resolution of inflammation, Nat. Rev. Immunol., 10, 427, 10.1038/nri2779

Song, 2005, PDGFRbeta+ perivascular progenitor cells in tumours regulate pericyte differentiation and vascular survival, Nat. Cell Biol., 7, 870, 10.1038/ncb1288

Song, 2009, Overexpression of platelet-derived growth factor-BB increases tumor pericyte content via stromal-derived factor-1alpha/CXCR4 axis, Cancer Res., 69, 6057, 10.1158/0008-5472.CAN-08-2007

Soriano, 1994, Abnormal kidney development and hematological disorders in PDGF beta-receptor mutant mice, Genes Dev., 8, 1888, 10.1101/gad.8.16.1888

Stagg, 2011, CD73-deficient mice have increased antitumor immunity and are resistant to experimental metastasis, Cancer Res., 71, 2892, 10.1158/0008-5472.CAN-10-4246

Stenzel, 2009, Peripheral mural cell recruitment requires cell-autonomous heparan sulfate, Blood, 114, 915, 10.1182/blood-2008-10-186239

Stout, 1942, Hemangiopericytoma: a vascular tumor featuring Zimmermann's pericytes, Ann. Surg., 116, 26, 10.1097/00000658-194207000-00004

Stratman, 2009, Pericyte recruitment during vasculogenic tube assembly stimulates endothelial basement membrane matrix formation, Blood, 114, 5091, 10.1182/blood-2009-05-222364

Stratman, 2010, Endothelial-derived PDGF-BB and HB-EGF coordinately regulate pericyte recruitment during vasculogenic tube assembly and stabilization, Blood, 116, 4720, 10.1182/blood-2010-05-286872

Stratman, 2011, VEGF and FGF prime vascular tube morphogenesis and sprouting directed by hematopoietic stem cell cytokines, Blood, 117, 3709, 10.1182/blood-2010-11-316752

Sundberg, 2002, Stable expression of angiopoietin-1 and other markers by cultured pericytes: phenotypic similarities to a subpopulation of cells in maturing vessels during later stages of angiogenesis in vivo, Lab. Invest., 82, 387, 10.1038/labinvest.3780433

Suri, 1996, Requisite role of angiopoietin-1, a ligand for the TIE2 receptor, during embryonic angiogenesis, Cell, 87, 1171, 10.1016/S0092-8674(00)81813-9

Tachibana, 2005, Selective role of a distinct tyrosine residue on Tie2 in heart development and early hematopoiesis, Mol. Cell. Biol., 25, 4693, 10.1128/MCB.25.11.4693-4702.2005

Takakura, 2000, A role for hematopoietic stem cells in promoting angiogenesis, Cell, 102, 199, 10.1016/S0092-8674(00)00025-8

Tang, 2008, White fat progenitor cells reside in the adipose vasculature, Science, 322, 583, 10.1126/science.1156232

Tavazoie, 2008, A specialized vascular niche for adult neural stem cells, Cell Stem Cell, 3, 279, 10.1016/j.stem.2008.07.025

Temmesfeld-Wollbrück, 2007, Adrenomedullin and endothelial barrier function, Thromb. Haemost., 98, 944, 10.1160/TH07-02-0128

Thurston, 1999, Leakage-resistant blood vessels in mice transgenically overexpressing angiopoietin-1, Science, 286, 2511, 10.1126/science.286.5449.2511

Thurston, 2005, Angiopoietin 1 causes vessel enlargement, without angiogenic sprouting, during a critical developmental period, Development, 132, 3317, 10.1242/dev.01888

Tidhar, 2001, A novel transgenic marker for migrating limb muscle precursors and for vascular smooth muscle cells, Dev. Dyn., 220, 60, 10.1002/1097-0177(2000)9999:9999<::AID-DVDY1089>3.0.CO;2-X

Tilton, 1979, Pericyte-endothelial relationships in cardiac and skeletal muscle capillaries, Microvasc. Res., 18, 325, 10.1016/0026-2862(79)90041-4

Uemura, 2002, Recombinant angiopoietin-1 restores higher-order architecture of growing blood vessels in mice in the absence of mural cells, J. Clin. Invest., 110, 1619, 10.1172/JCI0215621

Urness, 2000, Arteriovenous malformations in mice lacking activin receptor-like kinase-1, Nat. Genet., 26, 328, 10.1038/81634

Varga, 2008, Fibrosis in systemic sclerosis, Rheum. Dis. Clin. North Am., 34, 115, 10.1016/j.rdc.2007.11.002

Vates, 2010, Pericyte constriction after stroke: the jury is still out, Nat. Med., 16, 959, 10.1038/nm0910-959

Vikkula, 1996, Vascular dysmorphogenesis caused by an activating mutation in the receptor tyrosine kinase TIE2, Cell, 87, 1181, 10.1016/S0092-8674(00)81814-0

Voisin, 2010, Venular basement membranes ubiquitously express matrix protein low-expression regions: characterization in multiple tissues and remodeling during inflammation, Am. J. Pathol., 176, 482, 10.2353/ajpath.2010.090510

Vokes, 2004, Hedgehog signaling is essential for endothelial tube formation during vasculogenesis, Development, 131, 4371, 10.1242/dev.01304

Wang, 2009, Regulation of vascular permeability by sphingosine 1-phosphate, Microvasc. Res., 77, 39, 10.1016/j.mvr.2008.09.005

Wang, 2006, Venular basement membranes contain specific matrix protein low expression regions that act as exit points for emigrating neutrophils, J. Exp. Med., 203, 1519, 10.1084/jem.20051210

Wei, 2011, Fibrosis in systemic sclerosis: emerging concepts and implications for targeted therapy, Autoimmun. Rev., 10, 267, 10.1016/j.autrev.2010.09.015

Weskamp, 2010, Pathological neovascularization is reduced by inactivation of ADAM17 in endothelial cells but not in pericytes, Circ. Res., 106, 932, 10.1161/CIRCRESAHA.109.207415

Wessels, 2004, The epicardium and epicardially derived cells (EPDCs) as cardiac stem cells, Anat. Rec. A Discov. Mol. Cell. Evol. Biol., 276, 43, 10.1002/ar.a.10129

Wilm, 2005, The serosal mesothelium is a major source of smooth muscle cells of the gut vasculature, Development, 132, 5317, 10.1242/dev.02141

Winkler, 2004, Kinetics of vascular normalization by VEGFR2 blockade governs brain tumor response to radiation: role of oxygenation, angiopoietin-1, and matrix metalloproteinases, Cancer Cell, 6, 553

Winkler, 2010, Pericyte-specific expression of PDGF beta receptor in mouse models with normal and deficient PDGF beta receptor signaling, Mol. Neurodegener., 5, 32, 10.1186/1750-1326-5-32

Wipff, 2008, Integrins and the activation of latent transforming growth factor beta1 - an intimate relationship, Eur. J. Cell Biol., 87, 601, 10.1016/j.ejcb.2008.01.012

Xian, 2006, Pericytes limit tumor cell metastasis, J. Clin. Invest., 116, 642, 10.1172/JCI25705

Yang, 1999, Angiogenesis defects and mesenchymal apoptosis in mice lacking SMAD5, Development, 126, 1571, 10.1242/dev.126.8.1571

Yemisci, 2009, Pericyte contraction induced by oxidative-nitrative stress impairs capillary reflow despite successful opening of an occluded cerebral artery, Nat. Med., 15, 1031, 10.1038/nm.2022

Yonenaga, 2005, Absence of smooth muscle actin-positive pericyte coverage of tumor vessels correlates with hematogenous metastasis and prognosis of colorectal cancer patients, Oncology, 69, 159, 10.1159/000087840

Yu, 2010, Heparin-binding EGF-like growth factor protects pericytes from injury, J. Surg. Res.

Zachariah, 2010, Neural crest-derived pericytes promote egress of mature thymocytes at the corticomedullary junction, Science, 328, 1129, 10.1126/science.1188222

Zhou, 2008, Epicardial progenitors contribute to the cardiomyocyte lineage in the developing heart, Nature, 454, 109, 10.1038/nature07060

Zhu, 2008, NG2 cells generate both oligodendrocytes and gray matter astrocytes, Development, 135, 145, 10.1242/dev.004895

Zimmermann, 1923, Der feinere bau der blutcapillares, Z. Anat. Entwicklungsgesch., 68, 10.1007/BF02593544

Zlokovic, 2008, The blood-brain barrier in health and chronic neurodegenerative disorders, Neuron, 57, 178, 10.1016/j.neuron.2008.01.003

Zozulya, 2008, Pericyte-endothelial cell interaction increases MMP-9 secretion at the blood-brain barrier in vitro, Brain Res., 1189, 1, 10.1016/j.brainres.2007.10.099