Origin, causes and effects of increased nitrite concentrations in aquatic environments

Sarah Philips1, Hendrikus J. Laanbroek2, Willy Verstraete1
1Laboratory of Microbial Ecology and Technology, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium
2Centre for Limnology, Netherlands Institute of Ecology, Rijksstraatweg 6, 3631 AC Nieuwersluis, Netherlands

Tóm tắt

Từ khóa


Tài liệu tham khảo

Abeling U & Seyfried CF (1993) Anaerobic-aerobic treatment of potato-starch Wastewater. Wat. Sci. Tech. 28(2): 165-176

Alcaraz G, Chiappa Carrara X & Vanegas C (1997) Temperature tolerance of Penaeus setiferus postlarvae exposed to ammonia and nitrite. Aquat. Toxicol. 39(3-4): 345-353

Alcaraz G & Espina S (1997) Scope for growth of juvenile grass carp Ctenopharyngodon idella exposed to nitrite. Comp. Biochem. Physiol. 116C(1): 85-88

Alleman JE & Irvine RL (1980) Nitrification in the sequencing batch biological reactor. J. WPCF. 52(11): 2747-2754

Alleman JE (1984) Elevated nitrite occurrence in biological wastewater treatment systems. Wat. Sci. Tech. 17: 409-419

Alleman J & Preston K (1991) The biology of nitrifying bacteria. In Second annual workshop on commercial aquaculture using water recirculating systems, Purdue University, West Lafayette, 15-16 November 1991

Almeida JS, JÚlio SM, Reis MAM & Carrondo MJT (1995a) Nitrite inhibition of denitrification by Pseudomonas fluorescens. Biotechnol. Bioeng. 46: 194-201

Almeida JS, Reis MAM & Carrondo MJT (1995b) Competition between nitrate and nitrite reduction in denitrification by Pseudomonas fluorescens. Biotechnol. Bioeng. 46(5): 476-484

Anthonisen AC, Loehr RC, Prakasam TBS & Srinath EG (1976) Inhibition of nitrification by ammonia and nitrous acid. J. WPCF. 48(5): 835-852

Arquiaga MC, Canter LW & Sabatini DA (1993) Microbiology of high-sodium-nitrite-wastewater treatment. Environ. Pollut. 81: 1-6

Atlas RM & Bartha R (1993) Microbial Ecology-Fundamentals and Applications. The Benjamin/Cummings Publishing Company, Redwood City, California

Atwood HL, Fontenot QC, Tomasso JR and Isely JJ (2001) Toxicity of nitrite to Nile tilapia: Effect of fish size and environmental chloride. N. Am. J. Aquacult. 63(1): 49-51

Balmelle B, Nguyen KM, Capdeville B, Cornier JC & Deguin A (1992) Study of factors controlling nitrite build-up in biological processes for water nitrification. Wat. Sci. Tech., 26(5-6): 1017-1025

Barak Y, Tal Y & Van Rijn J (1998) Light-mediated nitrite accumulation during denitrification by Pseudomonas sp. strain JR12. Appl. Environ. Microb. 64: 813-817

Barak Y, Tal Y & Van Rijn J (2000) Relationship between nitrite reduction and active phosphate uptake in the phosphateaccumulating dentrifier Pseudomonas sp. strain JR 12. Appl. Environ. Microb. 66(12): 5236-5240

Barnes D & Bliss PJ (1983) Biological Control of Nitrogen in Wastewater Treatment. E. & F.N. Spon, New York, 146 pp

Baumann B, Snozzi M, Van der Meer JR & Zehder AJB (1997) Development of stable denitrifying cultures during repeated aerobic-anaerobic transient periods. Water Res. 31: 1947-1954

Bilski P, Szychlinski J & Oleksy E (1988) Oxygen consumption in two aqueous systems: excited nitrite ions-oxygen and nitrite ions-singlet oxygen. J. Photochem. Photobiol. A: Chem 45: 269-288

Blackall LL (2000) A summary of recent microbial discoveries in biological nutrient removal from wastewater. Aust. Biotech. 10(3): 29-31

Bock E (1965) Vergleichende Untersuchung über die Wirkung sichtbaren Lichtes auf Nitrosomonas europaea und Nitrobacter winogradskyi. Arch. Microbiol. 51: 18-41

Bock E (1978) Lithotrophic and chemoorganotrophic growth of nitrifying bacteria. In: Schlessinger D (Ed), Microbiology (pp. 310-314). American Society of Microbiology, Washington, DC

Bock E, Koops HP & Harms H (1986) Cell biology of nitrifying bacteria. In: Prosser JI (Ed), Nitrification (pp 17-38). IRL Press, Washington, DC

Bock E, Koops HP & Harms H (1989) Nitrifying bacteria. In: HG Schlegel & B Bowien (Eds), Autotrophic Bacteria (pp 81-96). Science Tech Publishers, Madison, Wisconsin

Bock E, Koops HP, Harms H & Ahlers B (1991) The biochemistry of nitrifying organisms. In: JM Shively & LL Barton (Eds), Variations in Autotrophic Life. Academic Press, New York

Bonin P (1996) Anaerobic nitrate reduction to ammonium in two strains isolated from costal marine sediment: A dissimilatory pathway. FEMS Microbiol. Ecol. 19(1): 27-38

Bovendeur J (1989) Fixed-biofilm reactors applied to waste water treatment and aquacultural water recirculating systems. Ph.D. Thesis, Agricultural University Wageningen, The Netherlands

Bradberry SM, Gazzard B and Vale JA (1994) Methemoglobinemia caused by the accidental contamination of drinking water with sodium nitrite. J. Toxicol-Clin. Toxic. 32: 173-178

Bredt DS (1999) Endogenous nitric oxide synthesis: Biological functions and pathophysiology. Free Radical Res. 31(6): 577-596

Brown & Caldwell (1975) Process design manual for nitrogen control. U.S. Environ. Prot. Agency Technology Transfer, EPA/625/1-75-007, PB-259 149/3BE

Brunning-Fann CS & Kaneene JB (1993a) The effects of nitrate, nitrite, and N-nitroso compounds on animal health. Vet. Hum. Toxicol. 35(3): 237-253

Brunning-Fann CS & Kaneene JB (1993b) The effects of nitrate, nitrite, and N-nitroso compounds on human health. Vet. Hum. Toxicol. 35(6): 521-538

Burrell PC, Keller J & Blackall L (1998) Microbiology of a nitriteoxidizing Bioreactor. Appl. Environ. Microb. 64: 1878-1883

Burrell P, Keller J & Blackall L (1999) Characterisation of the bacterial consortium involved in nitrite oxidation in activated sludge. Wat. Sci. Tech. 39(6): 45-52.

Casey TG, Wentzel MC & Ekama GA (1999a) Filamentous organism bulking in nutrient removal activated sludge systems. Paper 10: Metabolic behaviour of heterotrophic facultative aerobic organisms under aerated/unaerated conditions. Water SA. 25(4): 425-442

Casey TG, Wentzel MC & Ekama GA (1999b) Filamentous organism bulking in nutrient removal activated sludge systems. Paper 11: A biochemical/microbiological model for proliferation of anoxic-aerobic (AA) filamentous organisms. Water SA. 25(4): 443-451

Casey TG, Wentzel MC, Ekama GA, Loewenthal RE & Marais GvR (1994) A hypothesis for the causes and control of anoxicaerobic (AA) filament bulking in nutrient removal activated sludge systems. Wat. Sci. Tech. 29(7): 203-212

Caskey WH & Tiedje JM (1980) The reduction of nitrate to ammonium by a Clostridium sp. isolated from soil. J. Gen. Microbiol. 119: 217-223

Castignetti D & Gunner HB (1982) Differential tolerance of hydroxylamine by an Alcaligenes sp., a heterotrophic nitrifier, and by Nitrobacter agilis. Can. J. Microbiol. 28: 148-150

Ceçen F & Gönenç IE (1995) Criteria for nitrification and denitrification of high-strength wastes in 2 upflow submerged filters. Water Environ. Res. 67(2): 132-142

Chen JC & Chin TS (1988) Acute toxicity of nitrite to tiger prawn, Penaeus monodon, larvae. Aquaculture 69: 253-262

Cheng SY & Chen JC (1998) Effects of nitrite exposure on the hemolymph electrolyte, respiratory protein and free amino acid levels and water content of Penaeus japonicus. Aquat. Toxicol. 44: 129-139

Cheng SY & Chen JC (2000) Accumulation of nitrite in the tissues of Penaeus monodon exposed to elevated ambient nitrite after different time periods. Arch. Environ. Con. Tox. 39(2): 183-192

Cole J (1996) Nitrate reduction to ammonia by enteric bacteria: Redundancy, or a strategy for survival during oxygen starvation? FEMS Microbiol. Lett. 136: 1-11

Cui X, Joannou CL, Hughes MN & Cammack R (1992) The bactericidal effects of transition metal complexes containing the NO+ group of the food-spoilage bacterium Clostridium sporogenes. FEMS Microbiol. Lett. 98: 67-70

Dahl C, Sund C, Kristensen GH & Vredenbregt L (1997) Combined biological nitrification and denitrification of highsalinity wastewater Wat. Sci. Tech. 36(2-3): 345-352

Daims H, Nielsen PH, Nielsen JL, Juretschko S & Wagner M (2000) Novel Nitrospira-like bacteria as dominant nitrite-oxidizers in biofilms from wastewater treatment plants: diversity and in situ physiology. Wat. Sci. Tech. 41(4-5): 85-90

De Boer W, Klein Gunnewiek PJA, Veenhuis M, Bock E & Laanbroek HJ (1991) Nitrification at low pH by aggregated chemolithotrophic bacteria. Appl. Environ. Microbiol. 57(12): 3600-3604

Dinçer AR & Kargi F (1999) Salt inhibition of nitrification and denitrification in saline wastewater. Environ. Technol. 20: 1147-1153

Doblander C & Lackner R (1997) Oxidation of nitrite to nitrate in isolated erythrocytes: A possible mechanism for adaptation to environmental nitrite. Can. J. Fish. Aquat. Sci. 54(1): 157-161

Drysdale GD, Kasan HC & Bux F (2001) Assessment of denitri-fication by the ordinary heterotrophic organisms in an NDBEPR activated sludge system. Wat. Sci. Tech. 43(1): 147-154

Eddy FB & Williams EM (1994) Freshwater fish and nitrite. In Howells G (Ed), Water Quality for Freshwater Fish (pp 117-143). Gordon and Beach Science Publ., Switzerland

Eilersen AM, Henze M & Kloft L (1995) Effect of volatile fattyacids and trimethylamine on denitrification in activated sludge. Water Res. 29(5): 1259-1266

Eilersen AM, Henze M & Kloft L (1994) Effect of volatile fatty-acids and trimethylamine on nitrification in activated sludge Water Res. 28(6): 1329-1336

European Council Directive (1998) dd. 3/11/1998 concerning the quality of water destined for human consumption. Reference number JOLI 1998/330-4EN

European Economic Community (1978) A directive of the quality of freshwater needing protection or improvement in order to support fish life. Off. J. Eur. Comm. L222: 35-54

Fdz-Polanco F, Villaverde S & Garcia PA (1994) Temperature effect on nitrifying bacteria activity in biofilters: activation and free ammonia inhibition. Wat. Sci. Tech. 30(11): 121-130

Fdz-Polanco F, Villaverde S & Garcia PA (1996) Effects causing nitrite build-up in biofilters. Wat. Sci. Tech. 34(3): 371-378

Ferguson SJ (1994) Denitrification and its control. Ant. Leeuw. Int. J. G. 66: 89-110

Focht D & Verstraete W (1977) Biochemical ecology of nitrification and denitrification. In Alexander M (Ed) Advances in Microbial Ecology, vol. 1 (pp 135-214). Plenum Press, New York

Fontenot QC & Isely JJ (1998) Toxicity of ammonia and nitrite to Shortnose sturgeon. Meeting of the Southern Division of the American Fisheries Society Midyear Meeting, Lexington, Kentucky

Frances J, Allan GL & Nowak BF (1998) The effects of nitrite on the short-term growth of silver perch (Bidyanus bidyanus). Aquaculture 163(1-2): 63-72

Furumai H, Tagui H & Fujita K (1996) Effects of pH and alkalinity on sulfur-denitrification in a biological granular filter. Wat. Sci. Tech. 34(1-2): 355-362

item Gelda RK, Effler SW & Brooks CM (1999) Nitrite and the two stages of nitrification in nitrogen polluted Onondaga lake, New York. J. Environ. Qual. 28: 1505-1517

Gerards S, Duyts H & Laanbroek HJ (1998) Ammonium-induced inhibition of ammonium-starved Nitrosomonas europaea cells in soil and sand slurries. FEMS Microbiol. Ecol. 26: 269-280

Gerber JM (1997) Nutrition and migraine: Review and recommended strategies JNMS-J. Neuromusc. Sys. 5(3): 87-94

Ginestet P, Audic J-M, Urbain V & Block J-C (1998) Estimation of nitrifying bacterial activities by measuring oxygen uptake in the presence of the metabolic inhibitors allylthiourea and azide. Appl. Environ. Microb. 64(6): 2266-2268

Glass C & Silverstein J (1998) Denitrification kinetics of high nitrate concentration water: pH effect on inhibition and nitrite accumulation. Water Res. 32(3): 831-839

Glass C, Silverstein J & Oh J (1997) Inhibition of denitrification in activated sludge by nitrite Water Environ. Res. 69(6): 1086-1093

Gomez MA, Gonzalez-Lopez J & Hontoria-Garcia E (2000) Influence of carbon source on nitrate removal of contaminated groundwater in a denitrifying submerged filter. J. Haz. Mat. B80: 69-80

Gonzales JF, Dal Valle PL, Thohan S & Kane AS (2000) Effects of waterborne nitrite on phase I-II biotransformation in channel catfish (Ictalurus punctatus). Mar. Environ. Res. 50(1-5): 29-32

Goreau TJ, Kaplan WA, Wofsy SC, McElroy MB, Valois FW & Watson SW (1980) Production of NO2- and N2O by nitrifying bacteria at reduced concentrations of oxygen. Appl. Environ. Microb. 40: 526-532

Groeneweg J, Sellner B & Tappe W (1994) Ammonia oxidation in Nitrosomonas at NH3 concentrations near Km: Effects of pH and temperature. Water Res. 28: 2561-2566

Grosell M & Jensen FB (2000) Uptake and effects of nitrite in the marine teleost fish Platichthys flesus. Aquat. Toxicol. 50(1-2): 97-107

Grundmann GL & Debouzie D (2000) Geostatistical analysis of the distribution of NH4+ and NO2--oxidizing bacteria and serotypes at the millimeter scale along a soil transect. FEMS Microbiol. Ecol. 34: 57-62

Guerrero MA & Jones RD (1996) Photoinhibition of marine nitrifying bacteria. 1. Wavelength-dependent response. Mar. Ecol.-Prog. Ser. 141(1-3): 183-192

Hanaki K, Chalermraj W & Shinichiro O (1990a) Effects of the activity of heterotrophs on nitrification in a suspended-growth reactor. Water Res. 24(3): 289-296

Hanaki K, Chalermraj W & Shinichiro O (1990b) Nitrification at low levels of dissolved oxygen with and without organic loading in a suspended-growth reactor. Water Res. 24(3): 297-302

Hao OJ & Chen JM (1994) Factors affecting nitrite buildup insubmerged filter system. J. Environ. Eng.-ASCE. 120(5): 1298-1307

Harada H, Ando H & Momonoi K (1987) Process analysis of fluidized-bed biofilm reactor for denitrification. Wat. Sci. Tech. 19(1-2): 151-162

Harris JO, Maguire GB & Handlinger JH (1998) Effects of chronic exposure of greenlip abalone, Haliotis laevigata Donovan, to high ammonia, nitrite, and low dissolved oxygen concentrations on gill and kidney structure. J. Shellfish Res. 17(3): 683-687

He YJ & Bishop PL (1994) Effect of acid orange 7 on nitrification process. J. Environ. Eng.-New York, 120(1): 108-121

Hellinga C, Van Loosdrecht MCM & Heijnen JJ (1997) The Sharon process for nitrogen removal in ammonium rich wastewater. Med. Fac. Landbouww. Univ. Gent. 62: 1743-1750

Henze M, Grady CPL Jr, Gujer W, Marais GVR & Matsuo T (1987) Activated sludge model No. 1. IAWPRC Scientific and Technical Reports No. 1, London, UK

Hiorns WD, Hastings RC, Head IM, McCarthy AJ, Saunders JR, Pickup RW & Hall GH (1995) Amplification of 16S ribosomal RNA genes of autotrophic ammonia-oxidizing bacteria demonstrates the ubiquity of Nitrosospiras in the environment. Microbiology 141: 2793-2800

Hockenbury MR & Grady GPL (1977) Inhibition of nitrification-effects of selected organic compounds. J.WPCF. 49: 768-777

Holub W, Przytocka-Jusiak M, Blaszczyk M & Mycielski R (2000) Nitrite as agent selecting anaerobic phenol-degrading microflora in petroleum refining sediments. Water Res. 34(4): 1354-1358

Horan NJ & Azimi AA (1992) The effects of transient nitrogen loadings on nitrifying activated sludges in completely mixed and plug-flow reactors. Water Res 26(3): 279-284

Hovanec TA, Taylor LT, Blakis A & Delong EF (1998) Nitrospiralike bacteria associated with nitrite oxidation in freshwater aquaria. Appl. Environ. Microb. 64(1): 258-264

Hu SS (1990) Acute substrate-intermediate-product related inhibition of nitrifiers. M.S. Thesis, School of Civil Engineering, Purdue University, West Lafayette, Indiana, USA

Hunik JH, Tramper J & Wijffels RH (1994) A strategy to scale up nitrification processes with immobilized cells of Nitrosomonas europaea and Nitrobacter agilis. Bioprocess Eng. 11: 73-82

Jetten MSM, Logemann S, Muyzer G, Robertson LA, de Vries S, van Loosdrecht MCM & Kuenen JG (1997) Novel principles in the microbial conversion of nitrogen compounds. Ant. Leeuw. Int. J. G. 71: 75-93

Jetten MSM, Strous M, van de Pas-Schoonen KT, Schalk J, van Dongen UGJM, van de Graaf AA, Logemann S, Muyzer G, van Loosdrecht MCM & Kuenen JG (1999) The anaerobic oxidation of ammonium. FEMS Microbiol. Rev. 22: 421-437

Joo S-H, Kim D-J, Yoo I-K, Park K & Cha G-C (2000) Partial nitrification in an upflow biological aerated filter by O2 limitation. Biotechnol. Lett. 22: 937-940

Jooste SHJ & Vanleeuwen J (1993) Induction of nitrite build-up in water by some common disinfectants. Water SA. 19(2): 107-112

Juretschko S, Timmermann G, Schmid M, Schleifer K-H, Pommerening-Röser A, Koops H-P & Wagner M (1998) Combined molecular and conventional analyses of nitrifying bacterium diversity in activated sludge: Nitrososcoccus mobilis and Nitrospira-like bacteria as dominant Populations. Appl. Environ. Microb. 64(8): 3042-3051

Kamath S, Sabatini DA & Canter LW (1991) Biological nitrification/denitrification of high sodium nitrite (navy shipyard) wastewater. Environ. Pollut. 69(1): 25-38

Kamstra A, Span JA & van Weerd JH (1996) The acute toxicity and sub-lethal effects of nitrite on growth and feed utilization of European eel Anguilla anguilla (L.). Aqucault. Res. 27: 903-911

Kaplan D, Wilhelm R & Abeliovich A (2000) Interdependent environmental factors controlling nitrification in waters. Wat. Sci. Tech. 42(1-2): 167-172

Kelly DP (1978) Bioenergetics of chemolithotrophic bacteria. In: Bull AT and Meadow PM (Eds) Companion to Microbiology (pp 363-386). Longman, London

Kelso BHL, Smith RV & Laughlin RJ (1999) Effects of carbon substrates on nitrite accumulation in freshwater sediments. Appl. Environ. Microb. 65(1): 61-66

Kelso BHL, Smith RV, Laughlin RJ & Lennox SD (1997) Dissimilatory nitrate reduction in anaerobic sediments leading to river nitrite accumulation. Appl. Environ. Microb. 63: 4679-4685

Kieber RJ, Li A & Seaton PJ (1999) Production of nitrite from the photodegradation of dissolved organic matter in natural waters. Environ. Sci. Technol. 33(7): 993-998

Kielemoes J, De Boever P & Verstraete W (2000) Influence of denitrification on the corrosion of iron and stainless steel powder. Environ. Sci. Technol. 34(4): 663-671

Kleiner D (1985) Bacterial ammonium transport. FEMS Microbiol. Rev. 38: 87-100

Klüber HD & Conrad R (1998a) Inhibitory effects of nitrate, nitrite, NO and N2O on methanogenesis and other redox processes in anoxic rice field soil. FEMS Microbiol. Ecol. 25: 301-318

Klüber HD & Conrad R (1998b) Inhibitory effects of nitrate, nitrite, NO and N2O on methanogenesis by Methanosarcina barkeri and Methanobacterium bryantii. FEMS Microbiol. Ecol. 25: 331-339

Knowles G, Downing AL & Barett MJ (1965) Determination of kinetic constants for nitrifying bacteria in mixed culture, with the aid of an electronic computer. J. Gen. Microbiol. 38: 263-278

Koch G & Siegrist H (1997) Denitrification with methanol in tertiary filtration at wastewater treatment plant Zürich-Werdhölzli. Wat. Sci. Tech. 36(1): 165-172

Kone S & Behrens U (1981) Zur Kinetik der Denitrifikation, teil 1: Mischpopulation und Acetat als Kohlenstoffquele (in German). Acta Hydrochim. Hydrobiol. 9: 523-533

Kornaros M & Lyberatos G (1997) Kinetics of aerobic growth of a denitrifying bacterium, Pseudomonas denitrificans in the presence of nitrates and/or nitrites. Water Res. 31: 479-488

Kornaros M, Zafiri C & Lyberatos G (1996) Kinetics of denitri-fication by Pseudomonas denitrificans under growth conditions limited by carbon and/or nitrate or nitrite. Water Environ. Res. 68: 934-945

Krul JM & Veeningen R (1977) The synthesis of the dissimilatory nitrate reductase under aerobic conditions in a number of denitrifying bacteria, isolated from activated sludge and drinking water. Water Res. 11: 39-43

Kuai L & Verstraete W (1998) Ammonium removal by the oxygen limited autotrophic nitrificaton and denitrification (OLAND) system. Appl. Environ. Microb. 64(11): 4500-4506

Kuba T, van Loosdrecht MCM & Heijnen JJ (1996) Effect of cyclic oxygen exposure on the activity of denitrifying phosphorus removing bacteria. Wat. Sci. Tech. 34(1-2): 33-40

Kuhlen R, Busch T, Max M, Reyle-Hahn M, Falke KJ & Rossaint R (1999) Fluctuations of inspired concentrations of nitric oxide and nitrogen dioxide during mechanical ventilation. Crit. Care. 3(1): 1-6

Laanbroek HJ, Bodelier PLE & Gerards S (1994) Oxygen consumption kinetics of Nitrosomonas europaea and Nitrobacter hamburgensis grown in mixed cultures at different oxygen concentrations. Arch. Microbiol. 161: 156-162

Laanbroek HJ & Gerards S (1993) Competition for limiting amounts of oxygen between Nitrosomonas europaea and Nitrobacter winogradsky grown in mixed continuous cultures. Arch. Microbiol. 159: 453-459

Laanbroek HJ & Woldendorp JW (1995) Activity of chemolithotrophic nitrifying bacteria under stress in natural soils. In: Jones JG (Ed) Advances in Microbial Ecology, vol. 14 (pp 275-304). Plenum Press, New York

Lee YW, Ong SK & Sato C (1997) Effects of heavy metals on nitrifying bacteria. Wat. Sci. Tech. 36(12): 69-74

Leu HG, Lee CD, Ouyang CF & Tseng HT (1998) Effects of organic matter on the conversion rates of nitrogenous compounds in a channel reactor under various flow conditions. Water Res. 32(3): 891-899

Lewis WM Jr. & Morris DP (1986) Toxicity of nitrite to fish: a review. T. Am. Fish. Soc. 115: 183-199

Mallick N, Mohn FH & Soeder CJ (2000) Evidence supporting nitrite-dependent NO release by the green microalga Scenedesmus obliquus. J. Plant Physiol. 157(1): 40-46

Marco A & Blaustein AR (1999) The effects of nitrite on the behavior and metamorphosis in cascades frogs (Rana cascadae). Environ. Toxicol. Chem. 18(5): 946-949

Marco A, Quilchano C & Blaustein AR (1999) Sensitivity to nitrate and nitrite in pond-breeding amphibians from the Pacific northwest, USA. Environ. Toxicol. Chem. 18(12): 2836-2839

Martienssen M & Schops R (1997) Biological treatment of leachate from solid waste landfill sites-alterations in the bacterial community during the denitrification process. Water Res. 31: 1164-1170

Martienssen M & Schops R (1999) Population dynamics of denitrifying bacteria in a model biocommunity. Water Res. 33(3): 639-646

Martienssen M, Simon J & Schulz R (1995) Capacities and limits of three different technologies for the biological treatment of leachate from solid waste landfill sites. Acta Biotechnol. 15: 269-276

Mauret M, Paul E, Puech-Costes E, Maurette MT & Baptiste P (1996) Application of experimental research methodology to the study of nitrification in mixed culture. Wat. Sci. Tech. 34(1): 245-252

Meade ME & Watts SA (1995) Toxicity of ammonia, nitrite and nitrate to juvenile Australian crayfish, Cherax quadricarnatus'. J. Shellfish Res. 14(2): 341-346

Meinhold J, Arnold E & Isaacs S (1999) Effect of nitrite on anoxic phosphate uptake in biological phosphorus removal activated sludge. Water Res. 33(8): 1871-1883

Merkel W, Bohler E & Frimmel FH (1993) Heterotrophic denitrification-The influence of oxygen and pH value. Acta Hydroch. Hydrob. 21(5): 249-257

Mobarry BK, Wagner M, Urbain V, Rittmann BE & Stahl DA (1996) Phylogenetic probes for analyzing abundance and spatial organization of nitrifying bacteria. Appl. Environ. Microb. 62(6): 2156-2162

Morgenroth E, Obermayer A, Arnold E, Bruhl A, Wagner M & Wilderer PA (2000) Effect of long-term idle periods on the performance of sequencing batch reactors. Wat. Sci. Tech. 41(1): 105-113

Mulder JW & van Kempen R (1997) N-removal by Sharon. Water Qual. Int. 15: 30-31

Muller EB, Stouthamer AH & van Verseveld HW (1995) A novel method to determine maximal nitrification rates by sewage sludge at a non-inhibitory nitrite concentration applied to determine maximal rates as a function of the nitrogen load. Water Res. 29(4): 1191-1197

Münch EV, Lant P & Keller J (1996) Simultaneous nitrification and denitrification in becnh-scale sequencing batch reactors. Water Res. 30: 277-284

Musvoto EV, Lakay MT, Casey TG, Wentzel MC & Ekama GA (1999) Filamentous organism bulking in nutrient removal activated sludge systems-Paper 8: The effect of nitrate and nitrite. Water SA. 25(4): 397-407

Murray I, Parsons JW & Robinson K (1975) Interrelationships between nitrogen-balance, pH and dissolved oxygen in an oxidation ditch treating farm animal waste. Water Res. 9(1): 25-30

Neufeld R, Greenfield J & Rieder B (1986) Temperature, cyanide and phenolic nitrification inhibition. Water Res. 20(5): 633-642

Nijhof M & Klapwijk A (1995) Diffusional transport mechanisms and biofilm nitrification characteristics influencing nitrite levels in nitrifying trickling filter effluents. Water Res. 29(10): 2287-2292

Nowak O, Svardal K & Kroiss H (1996) The impact of phosphorus deficiency on nitrification-Case study of a biological pretreatment plant for rendering plant effluent. Wat. Sci. Tech. 34(1-2): 229-236

Nowak O, Svardal K & Schweighofer P (1994) The dynamic behaviour of nitrifying activated sludge systems influenced by inhibiting wastewater compounds. Proceedings of Modelling and Control of Activated Sludge Processes, Copenhagen, Denmark, 22-24 August 1994

Oh J & Silverstein J (1999a) Acetate limitation and nitrite accumulation during denitrification. J. Environ. Eng.-ASCE. 125(3): 234-242

Oh J & Silverstein J (1999b) Oxygen inhibition of activated sludge denitrification. Water Res. 33(8): 1925-1937

Okabe S, Oozawa Y, Hirata K & Watanabe Y (1996) Relationship between population dynamics of nitrifiers in biofilms and reactor performance at various C:N ratios. Water Res. 30(7): 1563-1577

Okabe S, Satoh H & Watanabe Y (1999) In situ analysis of nitrifying biofilms as determined by in city hybridization and the use of microelectrodes Appl. Environ. Microb. 65(7): 3182-3191

Olson RJ (1981) Differential photoinhibition of marine nitrifying bacteria: A possible mechanism for the formation of the primary nitrite maximum. J. Mar. Res. 39: 227-238

Pak D & Chang W (2000) Simultaneous removal of nitrogen and phosphorus in a two-biofilter system. Wat. Sci. Tech. 41(12): 101-106

Parra G & Yufera M (1999) Tolerance response to ammonia and nitrite exposure in larvae of two marine fish species (gilthead seabream Sparus aurata L. and Senegal sole Solea senegalensis Kaup). Aquac. Res. 30(11-12): 857-863

Payne WJ (1973) Reduction of nitrogenous oxides by microorganisms. Bacteriol. Rev. 37: 409-452

Peng DC, Bernet N, Delgenes JP & Moletta R (2000) Effects of oxygen supply methods on the performance of a sequencing batch reactor for high ammonium nitrification. Water Environ. Res. 72(2): 195-200

Percheron G, Bernet N & Moletta R (1999) Interactions between methanogenic and nitrate reducing bacteria during the anaerobic digestion of an industrial sulfate rich wastewater. FEMS Microbiol. Ecol. 29(4): 341-350

Philips S & Verstraete W (2000) Inhibition of activated sludge by nitrite in the presence of proteins or amino acids. Environ. Technol. 21: 1119-11125

Poughon L, Dussap CG & Gros JB (1999) Dynamic model of a nitrifying fixed bed column: Simulation of the biomass distribution of Nitrosomonas and Nitrobacter and of transient behaviour of the column. Bioprocess Eng. 20(3): 209-221

Prakasam TBS & Loehr RC (1972) Microbial nitrification and denitrification in concentrated wastes. Water Res. 6: 859-869

Pullium JK, Dillehay DL & Webb S (1999) High mortality in zebrafish (Danio rerio). Contemp. Top. lab. Anim. 38(3): 80-83

Purkhold U, Pommering-Röser A, Juretschko S, Schmid MC, Koops H-P & Wagner M (2000) Phylogney of all recognized species of ammonia oxidizers based on comparative 16S rRNA and amoA sequence analysis: Implications for molecular diversity surveys. Appl. Environ. Microb. 66(12): 5368-5382

Quevedo M, Guynot E & Muxi L (1996) Denitrifying potential of methanogenic Sludge. Biotechnol. Lett. 18(12): 1363-1368

Randall CW & Buth D (1984a) Nitrite build-up in an activated sludge resulting form combined temperature and toxicity effects. J. WPCF. 56: 1045-1049

Randall CW & Buth D (1984b) Nitrite build-up in activated sludge resulting from temperature effects. J. WPCF. 56: 1039-1044

Rittman BE, Laspidou CS, Flax J, Stahl DA, Urbain V, Harduin H, van der Waarde J, Geurkink B, Henssen MJC, Brouwer H, Klapwijk A & Wetterauw M(1999) Molecular and modeling analyses of the structure and function of nitrifying sludge. Wat. Sci. Tech. 39(1): 51-59

Rols JL, Mauret M, Rahmani H, Ngyen KM, Capdeville B, Cornier JC & Deguin A (1994) Population dynamics and nitrite build-up in activated sludge and biofilm processes for nitrogen removal. Wat. Sci. Tech. 29(7): 43-51

Rottenberg H (1990) Decoupling of oxidative phosphorylation and photo-phosphorylation. Biochim. Biophys. Acta 1018: 1-17

Sauter LJ & Alleman JE (1980) A streamlined approach to biological nitrogen removal. J. Environ. Eng.-ASCE.: 296-306

Schramm A, de Beer D, Gieseke A & Amann R (2000) Microenvironments and distribution of nitrifying bacteria in a membranebound biofilm. Environ. Microbiol. 2(6): 680-686

Schramm A, de Beer D, Wagner M & Amann R (1998) Identification and activities in situ of Nitrosospira and Nitrospira spp. as dominant populations in a nitrifying fluidized bed reactor. Appl. Environ. Microb. 64(9): 3480-3485

Schuch R, Gensicke R, Merkel K & Winter J (2000) Nitrogen and DOC removal from wastewater streams of the metal-working industry. Water Res. 34(1): 295-303

Sharma B & Ahlert RC (1977) Nitrification and nitrogen removal. Water. Res. 11: 897-925

Sijbesma WFH, Almeida JS, Reis MAM & Santos H (1996) Uncoupling effect of nitrite during denitrification by Pseudomonas fluorescens: an in vivo 31P-NMR study. Biotechnol. Bioeng. 52: 176-182

Smith MS (1982) Dissimilatory reduction of NO2 - to NH4 + and N2O by a soil Citrobacter sp. Appl. Environ. Microb. 43(4): 854-860

Smith RV, Burns LC, Doyle RM, Lennox SD, Kelso BHL, Foy RH & Stevens RJ (1997a) Free ammonia inhibition of nitrification in river sediments leading to nitrite accumulation. J. Environ. Qual. 26: 1049-1055

Smith RV, Doyle RM, Burns LC & Stevens RJ (1997b) A model for nitrite accumulation in Soils. Soil Biol. Biochem. 29(8): 1241-1247

Smith RV, Foy RH, Lennox SD, Jordan C, Burns LC, Cooper JE & Stevens RJ (1995) Occurrence of nitrite in the Lough Neagh River System. J. Environ. Qual. 24: 952-959

Sørensen J (1978) Capacity for denitrification and reduction of nitrate to ammonia in a coastal marine sediment. Appl. Environ. Microb. 35(2): 301-305

Spector M (1998) Cocurrent biological nitrification and denitri-fication in wastewater treatment. Water Environ. Res. 70(7): 1242-1247

Stein LY & Arp DJ (1998) Loss of ammonia monooxygenase activity in Nitrosomonas europaea upon exposure to nitrite. Appl. Environ. Microb. 64(10): 4098-4102

Stein LY, Arp DJ & Hyman MR (1997) Regulation of the synthesis and activity of ammonia monooxygenase in Nitrosomonas europaea by altering pH to affect NH3 availability. Appl. Environ. Microb. 63(11): 4588-4592

Strous M (2000) Microbiology of anaerobic ammonium oxidation. Ph.D. Thesis, Technical University Delft, The Netherlands

Stüven R, Vollmer M & Bock E (1992) The impact of organic matter on nitric oxide formation by Nitrosomonas europaea. Arch. Microbiol. 158: 439-443

Surmacz-Gorska J, Cichon A & Miksch K (1997) Nitrogen removal from wastewater with high ammonia nitrogen concentration via shorter nitrification and denitrification. Wat. Sci. Tech. 36(10): 73-78

Surmacz-Gorska J, Gernaey K, Demuynck C, Vanrolleghem P & Verstraete W (1996) Nitrification monitoring in activated sludge by oxygen uptake rate (OUR) measurements. Water Res. 30: 1228-1236

Suthursan S & Ganczarczyk JJ (1986) Inhibition of nitrite oxidation during nitrification, some observations. Water Pollut. Res. J. Can. 21(2): 257-266

Suzuki I, Kwok SC & Dular U (1974) Ammonia or ammonium ion as substrate for oxidation by Nitrosomonas europaea cells and extracts. J. Bacteriol. 120: 556-558

Takai T, Hirata A, Yamauchi K & Inamori Y (1997) Effects of temperature and volatile fatty acids on nitrification denitrification activity in small scale anaerobic aerobic recirculation biofilm process. Wat. Sci. Tech. 35(6): 101-108

Tang NH, Blum DJW, Nirmalakhandan N & Speece RE (1992) QSAR parameters for toxicity of organic chemicals to Nitrobacter. J. Environ. Eng.-ASCE. 118(1): 17-37

Tappe W, Laverman A, Bohland M, Braster M, Rittershaus S, Groeneweg J & van Verseveld HW (1999) Maintenance energy demand and starvation recovery dynamics of Nitrosomonas europaea and Nitrobacter winogradsky cultivated in a retentostat with complete biomass retention. Appl. Environ. Microb. 65(6): 2471-2477

Tiedje JM, Sexstone AJ, Myrold DD & Robinson JA (1982) Denitrification: ecological niches, competition and survival. Ant. Leeuw. Int. J. G. 48: 569-583

Tomlinson TG, Boon AG & Trotman CNA (1966) Inhibition of nitrification in the activated sludge process of sewage disposal. J. Appl. Bacteriol. 29: 266-291

Tonkovic Z (1998) Nitrite accumulation at the Mornington sewage treatment plant-causes and significance (pp 165-172). 19th Biennial International Conference, Water Quality International 1998. IAWQ, Vancouver, Canada, 21-26 June 1998

Turk O & Mavinic DS (1986) Preliminary assessment of a shortcut in nitrogen removal from wastewater. Can. J. Civil Eng. 13: 600-605

Turk O & Mavinic DS (1989) Stability of nitrite build-up in an activated sludge system. J. WPCF. 61(8): 1440-1448

Van Leeuwen FXR (2000) Safe drinking water: The toxicologist's approach. Food Chem. Toxicol. 38: S51-S58

Van Rijn J, Tal Y & Barak Y (1996) Influence of volatile fatty acids on nitrite accumulation by a Pseudomonas stutzeri strain isolated from a denitrifying fluidized bed reactor. Environ. Microbiol. 62: 2615-2620

Vanzella A, Guerrero MA & Jones RD (1989) Effects of CO and light on ammonium and nitrite oxidation by chemolitotrophic bacteria. Mar. Ecol.-Prog. Ser. 57: 69-76

Venterea RT & Rolston DE (2000a) Mechanistic modeling of nitrite accumulation and nitrogen oxide gas emissions during nitrification. J. Environ. Qual. 29: 1741-1751

Venterea RT & Rolston DE (2000b) Mechanisms and kinetics of nitric and nitrous oxide production during nitrification in agricultural soil'. Glob. Change Biol. 6(3): 303-316

Verstraete W, Vanstaen H & Voets JP (1977) Adaptation to nitrification of activated sludge systems treating highly nitrogenous waters. J. WPCF. 49: 1604-1608

Verstraete W & Philips S (1998) Nitrification/denitrification processes and technologies in new contexts. Environ. Pollut. 102: 717-726

Villaverde S, Fdz-Polanco F & Garcia PA (1997) Influence of pH on nitrifying bacteria activity in submerged biofilters. Water Res. 31(5): 1180-1186

Villaverde S, Fdz-Polanco F & Garcia PA (2000) Nitrifying biofilm acclimation to free ammonia in submerged biofilters. Start-up influence. Water Res. 34(2): 602-610

Von Der Wiesche M & Wetzel A (1998) Temporal and spatial dynamics of nitrite accumulation in the river Lahn. Water Res. 32(5): 1653-1661

von Schulthess R, Kühni M & Gujer W (1995) Release of nitric and nitrous oxides from denitrifying activated sludge. Water Res. 29(1): 215-226

Wang W (1984) Time response of Nitrobacter to toxicity. Environ. Int. 10: 21-26

Ward BB (2000) Nitrification and the marine nitrogen cycle. In: Kirchman DL (Ed), Microbiol Ecology of the Oceans (pp 427-453). Wiley-Liss, New York

Weng YM, Hotchkiss JH & Babish JG (1992) N-nitrosamine and mutagenicity formation in Chinese salted fish after digestion. Food Addit. Contam. 9: 29-37

WHO (1996) Guidelines for drinking water quality (pp 940-949). Health criteria and other supporting information, World Health Organisation, Geneva

Wild D, Von Schulthess R & Gujer W (1995) Structured modelling of denitrification Intermediates. Wat. Sci. Tech. 31: 45-54

Wilderer PA, Jones WJ & Dau U (1987) Competition in denitrification systems affecting reduction rate and accumulation of nitrite. Water Res. 21: 239-245

Wise DJ & Tomasso JR (1989) Acute toxicity of nitrite to red drum, Sciaenops ocellatus: effect of salinity. J. World Aquacul. Soc. 20(4): 193-198

Wong-Chong GM & Loehr RC (1975) Kinetics of microbial nitrification. Water Res. 9(12): 1099-1106

Wong-Chong GM & Loehr RC (1978) Kinetics of microbial nitrification-Nitrite-nitrogen oxidation. Water Res. 12(8): 605-609

Wood PM (1986) Nitrification as a bacterial energy source. In: Prosser JI (Ed) Nitrification (pp 39-62). IRL Press, Oxford, UK

Wortman B & Wheaton F (1991) Temperature effects on biodrum nitrification. Aquacult. Eng. 10: 183-205

Yang L & Alleman JE (1992) Investigation of batchwise nitrite build-up by an enriched nitrification culture. Wat. Sci. Tech. 26(5-6): 997-1005

Zhang TC, Fu YC & Bishop PL (1995) Competition for substrate and space in Biofilms. Water Environ. Res. 67: 992-1003

Zhang SY, Wang JS, Jiang ZC & Chen MX (2000) Nitrite accumulation in an attapulgas clay biofilm reactor by fulvic acids. Bioresource Technol. 73(1): 91-93

Zumft WG (1993) The biological role of nitric oxide in bacteria. Arch. Microbiol. 160: 263-264

Zumft WG (1992) The denitrifying prokaryotes. In: Balows A, Trüper HG, Dworkin M, Harder W & Schleifer K-H (Eds), The Prokaryotes (pp 554-582). Springer-Verlag, Berlin