1Guangdong Key Laboratory for Innovative Development and Utilization of Forest Plant Germplasm, College of Forestry and Landscape Architectures, South China Agricultural University, Guangzhou, China
2State Key Laboratory for Conservation and Utilization of Subtropical Agro-Bioresources, Guangzhou, China
3College of Agriculture, South China Agricultural University, Guangzhou, China
4State Key Laboratory for Conservation and Utilization of Subtropical Agro-Bioresources, Guangxi Research Center for Microbial and Enzyme Engineering Technology, College of Life Science and Technology, Guangxi University, Nanning, China
5Guangdong Laboratory of Lingnan Modern Agriculture, Guangzhou, China