Harmful algal blooms: A climate change co-stressor in marine and freshwater ecosystems

Harmful Algae - Tập 91 - Trang 101590 - 2020
Andrew W. Griffith1,2, Christopher J. Gobler2
1Department of Biological Sciences, University of Southern California, Los Angeles, CA 90089, United States
2School of Marine and Atmospheric Sciences, Stony Brook University, Southampton, NY, 11968, United States

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Abele, 2002, Temperature-dependence of mitochondrial function and production of reactive oxygen species in the intertidal mud clam Mya arenaria, J. Exp. Biol., 205, 1831, 10.1242/jeb.205.13.1831

Anderson, 2012, HABs in a changing world: a perspective on harmful algal blooms, their impacts, and research and management in a dynamic era of climactic and environmental change, 3

Anderson, 1990, Dynamics and physiology of saxitoxin production by the dinoflagellates Alexandrium spp, Mar. Biol., 104, 511, 10.1007/BF01314358

Anderson, 2000

Anderson, 2002, Harmful algal blooms and eutrophication: nutrient sources, composition, and consequences, Estuaries, 25, 704, 10.1007/BF02804901

Asch, 2015, Climate change and decadal shifts in the phenology of larval fishes in the California Current ecosystem, Proc. Natl. Acad. Sci. U. S. A., 112, E4065, 10.1073/pnas.1421946112

Backer, 2003, Recreational exposure to aerosolized brevetoxins during Florida red tide events, Harmful Algae, 2, 19, 10.1016/S1568-9883(03)00005-2

Baden, 2005, Natural and derivative brevetoxins: historical background, multiplicity, and effects, Environ. Health Perspect., 113, 621, 10.1289/ehp.7499

Bauman, 2010, Tropical harmful algal blooms: an emerging threat to coral reef communities?, Mar. Pollut. Bull., 60, 2117, 10.1016/j.marpolbul.2010.08.015

Baumann, 2013, Decadal changes in the world’s coastal latitudinal temperature gradients, PLoS One, 8, 10.1371/journal.pone.0067596

Baumann, 2018, Quantifying metabolically driven pH and oxygen fluctuations in US nearshore habitats at diel to interannual time scales, Estuaries Coasts, 41, 1102, 10.1007/s12237-017-0321-3

Baumann, 2015, Large natural pH, CO2 and O2 fluctuations in a temperate tidal salt marsh on diel, seasonal, and interannual time scales, Estuaries Coasts, 38, 220, 10.1007/s12237-014-9800-y

Belkin, 2009, Rapid warming of large marine ecosystems, Prog. Oceanogr., 81, 207, 10.1016/j.pocean.2009.04.011

Braga, 2018, Combined effects of warming and acidification on accumulation and elimination dynamics of paralytic shellfish toxins in mussels Mytilus galloprovincialis, Environ. Res., 164, 647, 10.1016/j.envres.2018.03.045

Breitburg, 2018, Declining oxygen in the global ocean and coastal waters, Science, 359, 10.1126/science.aam7240

Bricelj, 1997, Aureococcus anophagefferens: causes and ecological consequences of brown tides in U.S. Mid-Atlantic coastal waters, Limnol. Oceanogr., 42, 1023, 10.4319/lo.1997.42.5_part_2.1023

Bricelj, 2007, Effects of brown tide (Aureococcus anophagefferens) on hard clam Mercenaria mercenaria larvae and implications for benthic recruitment, Mar. Ecol. Prog. Ser., 331, 147, 10.3354/meps331147

Bricelj, 2005, Sodium channel mutation leading to saxitoxin resistance in clams increases risk of PSP, Nature, 434, 763, 10.1038/nature03415

Brutemark, 2015, Growth, toxicity and oxidative stress of a cultured cyanobacterium (Dolichospermum sp.) under different CO2/pH and temperature conditions: environmental stress for cyanobacteria, Phycol. Res., 63, 56, 10.1111/pre.12075

Burford, 2019, Perspective: advancing the research agenda for improving understanding of cyanobacteria in a future of global change, Harmful Algae, 91

Burge, 2014, Climate change influences on marine infectious diseases: implications for management and society, Annu. Rev. Mar. Sci., 6, 249, 10.1146/annurev-marine-010213-135029

Burkholder, 2011, Bivalve shellfish aquaculture and eutrophication, 155

Burkholder, 2008, Mixotrophy, a major mode of nutrition for harmful algal species in eutrophic waters, Harmful Algae, 8, 77, 10.1016/j.hal.2008.08.010

Byrne, 2013, Multistressor impacts of warming and acidification of the ocean on marine invertebrates’ life histories, Integr. Comp. Biol., 53, 582, 10.1093/icb/ict049

Cai, 2011, Acidification of subsurface coastal waters enhanced by eutrophication, Nat. Geosci., 4, 766, 10.1038/ngeo1297

Cheng, 2007, Characterization of aerosols containing microcystin, Mar. Drugs, 5, 136, 10.3390/md504136

Colin, 2002, Latitudinal differentiation in the effects of the toxic dinoflagellate Alexandrium spp. on the feeding and reproduction of populations of the copepod Acartia hudsonica, Harmful Algae, 1, 113, 10.1016/S1568-9883(02)00007-0

Cosper, 1987, Recurrent and persistent brown tide blooms perturb coastal marine ecosystem, Estuaries, 10, 284, 10.2307/1351885

Cossellu, 2010, Recent environmental changes and filamentous algal mats in shallow bays on the Swedish west coast — a result of climate change?, J. Sea Res., 63, 202, 10.1016/j.seares.2010.01.004

Crain, 2008, Interactive and cumulative effects of multiple human stressors in marine systems, Ecol. Lett., 11, 1304, 10.1111/j.1461-0248.2008.01253.x

Davis, 2009, The effects of temperature and nutrients on the growth and dynamics of toxic and non-toxic strains of Microcystis during cyanobacteria blooms, Harmful Algae, 8, 715, 10.1016/j.hal.2009.02.004

DePasquale, 2015, Vulnerability of early life stage Northwest Atlantic forage fish to ocean acidification and low oxygen, Mar. Ecol. Prog. Ser., 523, 145, 10.3354/meps11142

Diaz, 2008, Spreading dead zones and consequences for marine ecosystems, Science, 321, 926, 10.1126/science.1156401

Doblin, 2006, Vertical migration of the toxic dinoflagellate Gymnodinium catenatum under different concentrations of nutrients and humic substances in culture, Harmful Algae, 5, 665, 10.1016/j.hal.2006.02.002

Donelson, 2011, Rapid transgenerational acclimation of a tropical reef fish to climate change, Nat. Clim. Change, 2, 30, 10.1038/nclimate1323

Doney, 2012, Climate change impacts on marine ecosystems, Annu. Rev. Mar. Sci., 4, 11, 10.1146/annurev-marine-041911-111611

Dong, 2014, Understanding strategy of nitrate and urea assimilation in a chinese strain of Aureococcus anophagefferens through RNA-Seq analysis, PLoS One, 9

Edwards, 2004, Impact of climate change on marine pelagic phenology and trophic mismatch, Nature, 430, 881, 10.1038/nature02808

Etheridge, 2005, Effects of temperature, irradiance, and salinity on photosynthesis, growth rates, total toxicity, and toxin composition for Alexandrium fundyense isolates from the Gulf of Maine and Bay of Fundy, Deep Sea Res. Part II Top. Stud. Oceanogr., 52, 2491, 10.1016/j.dsr2.2005.06.026

Flynn, 2018, Mixotrophy in harmful algal blooms: by whom, on whom, when, why, and what next, 113

Fraga, 1989, Chain-forming dinoflagellates: an adaptation to red tides, 281

Friedland, 2013, Thermal habitat constraints on zooplankton species associated with Atlantic cod (Gadus morhua) on the US Northeast Continental Shelf, Prog. Oceanogr., 116, 1, 10.1016/j.pocean.2013.05.011

Fu, 2008, A comparison of future increased CO2and temperature effects on sympatric Heterosigma akashiwo and Prorocentrum minimum, Harmful Algae, 7, 76, 10.1016/j.hal.2007.05.006

Fu, 2010, CO2 and phosphate availability control the toxicity of the harmful bloom dinoflagellate Karlodinium veneficum, Aquat. Microb. Ecol., 59, 55, 10.3354/ame01396

Fu, 2012, Global change and the future of harmful algal blooms in the ocean, Mar. Ecol. Prog. Ser., 470, 207, 10.3354/meps10047

Funkey, 2014, Hypoxia sustains cyanobacteria blooms in the Baltic sea, Environ. Sci. Tech., 48, 2598, 10.1021/es404395a

Galimany, 2017, Feeding behavior of eastern oysters Crassostrea virginica and hard clams Mercenaria mercenaria in shallow estuaries Marine, Eco. Prog Series, 567, 125, 10.3354/meps12050

Gilbert, 2010, Evidence for greater oxygen decline rates in the coastal ocean than in the open ocean, Biogeosciences, 7, 2283, 10.5194/bg-7-2283-2010

Glibert, 2006, The complex relationships between increases in fertilization of the earth, coastal eutrophication and proliferation of harmful algal blooms, 341

Glibert, 2014, Vulnerability of coastal ecosystems to changes in harmful algal bloom distribution in response to climate change: projections based on model analysis, Glob. Change Biol., 20, 3845, 10.1111/gcb.12662

Glibert, 2018, Nutrients and harmful algal blooms: dynamic kinetics and flexible nutrition, 93

Gobler, 2016, Hypoxia and acidification in ocean ecosystems: coupled dynamics and effects on marine life, Biol. Lett., 12, 10.1098/rsbl.2015.0976

Gobler, 2012, Ecosystem disruptive algal blooms of the brown tide species, Aureococcus anophagefferens and Aureoumbra lagunensis, Harmful Algae, 14, 36, 10.1016/j.hal.2011.10.013

Gobler, 2013, Short- and long-term consequences of larval stage exposure to constantly and ephemerally elevated carbon dioxide for marine bivalve populations, Biogeosciences, 10, 2241, 10.5194/bg-10-2241-2013

Gobler, 2005, A review of the causes, effects, and potential management of harmful brown tide blooms caused by Aureococcus anophagefferens (Hargraves et sieburth), Estuaries, 28, 726, 10.1007/BF02732911

Gobler, 2014, Hypoxia and acidification have additive and synergistic negative effects on the growth, survival, and metamorphosis of early life stage bivalves, PLoS One, 9, 10.1371/journal.pone.0083648

Gobler, 2017, Ocean warming since 1982 has expanded the niche of toxic algal blooms in the North Atlantic and North Pacific oceans, Proc. Natl. Acad. Sci., 114, 4975, 10.1073/pnas.1619575114

Gobler, 2018, Temperature, acidification, and food supply interact to negatively affect the growth and survival of the forage fish, Menidia beryllina (inland silverside), and Cyprinodon variegatus (sheepshead minnow), Front. Mar. Sci., 5, 10.3389/fmars.2018.00086

Green, 2009, Death by dissolution: sediment saturation state as a mortality factor for juvenile bivalves, Limnol. Oceanogr., 54, 1037, 10.4319/lo.2009.54.4.1037

Griffith, 2016, Temperature Controls the toxicity of the icthyotoxic dinoflagellate, Cochlodinium polykrikoides, Mar. Ecol. Prog. Ser., 545, 63, 10.3354/meps11590

Griffith, 2017, Transgenerational exposure of North Atlantic bivalves to ocean acidification renders offspring more vulnerable to low pH and additional stressors, Sci. Rep., 7, 10.1038/s41598-017-11442-3

Griffith, A.W., Doherty, O.M., Gobler, C.J., in press. Ocean warming along temperate western-boundaries of the Northern Hemisphere promotes an expansion of Cochlodinium polykrikoides blooms Proc. R. Soc. B.

Griffith, 2019, Differential mortality of North Atlantic bivalve molluscs during harmful algal blooms caused by the dinoflagellate,Cochlodinium (a.k.a. Margalefidinium) polykrikoides, Estuaries Coasts, 42, 190, 10.1007/s12237-018-0445-0

Guerrini, 2007, Influence of temperature, salinity and nutrient limitation on yessotoxin production and release by the dinoflagellate Protoceratium reticulatum in batch-cultures, Harmful Algae, 6, 707, 10.1016/j.hal.2007.02.006

Gupta, 2003, Comparative toxicity evaluation of cyanobacterial cyclic peptide toxin microcystin variants (LR, RR, YR) in mice, Toxicology, 188, 285, 10.1016/S0300-483X(03)00112-4

Hallegraeff, 2010, Ocean climate change, phytoplankton community responses, and harmful algal blooms: a formidable predictive challenge, J. Phycol., 46, 220, 10.1111/j.1529-8817.2010.00815.x

Halpern, 2008, A global map of human impact on marine ecosystems, Science, 319, 948, 10.1126/science.1149345

Hamasaki, 2001, Variability in toxicity of the dinoflagellate Alexandrium tamarense isolated from Hiroshima Bay, Western Japan, as a reflection of changing environmental conditions, J. Plankton Res., 23, 271, 10.1093/plankt/23.3.271

Harley, 2006, The impacts of climate change in coastal marine systems, Ecol. Lett., 9, 228, 10.1111/j.1461-0248.2005.00871.x

Harvey, 2013, Meta-analysis reveals complex marine biological responses to the interactive effects of ocean acidification and warming, Ecol. Evol., 3, 1016, 10.1002/ece3.516

Hattenrath-Lehmann, 2015, Nitrogenous nutrients promote the growth and toxicity of Dinophysis acuminata during estuarine bloom events, PLoS One, 10, 10.1371/journal.pone.0124148

Hattenrath-Lehmann, 2015, The effects of elevated CO2 on the growth and toxicity of field populations and cultures of the saxitoxin-producing dinoflagellate, Alexandrium fundyense, Limnol. Oceanogr., 60, 198, 10.1002/lno.10012

Hégaret, 2009, Biotoxin contamination and shellfish safety, Shellfish Saf. Qual., 43, 10.1533/9781845695576.1.43

Heisler, 2008, Eutrophication and harmful algal blooms: a scientific consensus, Harmful Algae, 8, 3, 10.1016/j.hal.2008.08.006

Helluy, 1991, Embryonic development of the American lobster (Homarus americanus): quantitative staging and characterization of an embryonic molt cycle, Biol. Bull., 180, 355, 10.2307/1542337

Hoagland, 2006, The economic effects of harmful algal blooms, 391

Hochachka, 2001, Mechanism, origin, and evolution of anoxia tolerance in animals, Comp. Biochem. Physiol. B, Biochem. Mol. Biol., 130, 435, 10.1016/S1096-4959(01)00408-0

Hoegh-Guldberg, 2010, The impact of climate change on the world’s marine ecosystems, Science, 328, 1523, 10.1126/science.1189930

Hoegh-Guldberg, 2014, The Ocean, 1655

Hu, 2015, Antioxidant responses of triangle sail mussel Hyriopsis cumingii exposed to harmful algae Microcystis aeruginosa and hypoxia, Chemosphere, 139, 541, 10.1016/j.chemosphere.2015.07.074

Hu, 2016, Combined effects of toxic cyanobacteria Microcystis aeruginosa and hypoxia on the physiological responses of triangle sail mussel Hyriopsis cumingii, J. Hazard. Mater., 306, 24, 10.1016/j.jhazmat.2015.11.052

IPCC, 2014, Climate change 2014: synthesis report

Jansson, 2015, Growth and survival in a changing environment: combined effects of moderate hypoxia and low pH on juvenile bivalve Macoma balthica, J. Sea Res., 102, 41, 10.1016/j.seares.2015.04.006

Kamiyama, 2010, Effect of temperature on production of okadaic acid, dinophysistoxin-1, and pectenotoxin-2 by Dinophysis acuminata in culture experiments, Aquat. Microb. Ecol., 60, 193, 10.3354/ame01419

Kennedy, 1982, Comparative gametogenic and spawning patterns of the oyster Crassostrea virginica in central Chesapeake Bay, USA, J. Shellfish Res., 2, 133

Kim, 1997, Recent harmful algal blooms and mitigation strategies in Korea, Ocean Polar Res., 19, 185

Kim, 1998, Harfmul algal bloom in Korean coastal waters focused on three fish-killing dinoflagellates

Kim, 2013, pH alters the swimming behaviors of the raphidophyte Heterosigma akashiwo: Implications for bloom formation in an acidified ocean, Harmful Algae, 26, 1, 10.1016/j.hal.2013.03.004

Kim, 2013, The effects of intermittent exposure to low-pH and low-oxygen conditions on survival and growth of juvenile red abalone, Biogeosciences, 10, 7255, 10.5194/bg-10-7255-2013

Kim, 2014, Combined effects of the cyanobacterial toxin microcystin-LR and environmental factors on life-history traits of indigenous cladoceran Moina macrocopa: combined toxicity of MC-LR and water environmental factors, Environ. Toxicol. Chem., 33, 2560, 10.1002/etc.2712

Kremp, 2012, Intraspecific variability in the response of bloom-forming marine microalgae to changed climate conditions: phenotypic variability and climate conditions, Ecol. Evol., 2, 1195, 10.1002/ece3.245

Kroeker, 2013, Impacts of ocean acidification on marine organisms: quantifying sensitivities and interaction with warming, Glob. Change Biol., 19, 1884, 10.1111/gcb.12179

Kudela, 2010, The role of nutrients in regulation and promotion of harmful algal blooms in upwelling systems, Prog. Oceanogr., 85, 122, 10.1016/j.pocean.2010.02.008

Landsberg, 2002, The effects of harmful algal blooms on aquatic organisms, Rev. Fish. Sci., 10, 113, 10.1080/20026491051695

Levin, 2015, Linking coasts and seas to address ocean deoxygenation, Nat. Clim. Change, 5, 401, 10.1038/nclimate2595

Litaker, 2008, Rapid enzyme-linked immunosorbent assay for detection of the algal toxin domoic acid, J.Shellfish Res., 27, 1301, 10.2983/0730-8000-27.5.1301

Lundholm, 1997, Studies on the marine planktonic diatom Pseudo-nitzschia. 2. Autecology of P. pseudodelicatissima based on isolates from Danish coastal waters, Phycologia, 36, 381, 10.2216/i0031-8884-36-5-381.1

Lundholm, 2004, Effect of pH on growth and domoic acid production by potentially toxic diatoms of the genera Pseudo-nitzschia and Nitzschia, Mar. Ecol. Prog. Ser., 273, 1, 10.3354/meps273001

Lürling, 2017, Eutrophication and warming boost cyanobacterial biomass and microcystins, Toxins, 9, 64, 10.3390/toxins9020064

McCabe, 2016, An unprecedented coastwide toxic algal bloom linked to anomalous ocean conditions, Geophys. Res. Lett., 43, 10366, 10.1002/2016GL070023

Mellado, 2018, Ocean acidification exacerbates the effects of paralytic shellfish toxins on the fitness of the edible mussel Mytilus chilensis, Sci. Total Environ.

Melzner, 2011, Food supply and seawater pCO2 impact calcification and internal shell dissolution in the blue mussel Mytilus edulis, PLoS One, 6, 10.1371/journal.pone.0024223

Michaelidis, 2005, Effects of long-term moderate hypercapnia on acid-base balance and growth rate in marine mussels Mytilus galloprovincialis, Mar. Ecol. Prog. Ser., 293, 109, 10.3354/meps293109

Miles, 2004, Isolation of pectenotoxin-2 from Dinophysis acuta and its conversion to pectenotoxin-2 seco acid, and preliminary assessment of their acute toxicities, Toxicon, 43, 1, 10.1016/j.toxicon.2003.10.003

Munday, 2013, Acute toxicities of saxitoxin, neosaxitoxin, decarbamoyl saxitoxin and gonyautoxins 1&4 and 2&3 to mice by various routes of administration, Toxicon, 76, 77, 10.1016/j.toxicon.2013.09.013

Naar, 2002, A competitive ELISA to detect brevetoxins from Karenia brevis (formerly Gymnodinium breve) in seawater, shellfish, and mammalian body fluid, Environ. Health Perspect, 110, 179, 10.1289/ehp.02110179

Noyes, 2009, The toxicology of climate change: environmental contaminants in a warming world, Environ. Int., 35, 971, 10.1016/j.envint.2009.02.006

Nye, 2009, Changing spatial distribution of fish stocks in relation to climate and population size on the Northeast United States continental shelf, Mar. Ecol. Prog. Ser., 393, 111, 10.3354/meps08220

O’Neil, 2012, The rise of harmful cyanobacteria blooms: the potential roles of eutrophication and climate change, Harmful Algae, 14, 313, 10.1016/j.hal.2011.10.027

Ogata, 1987, Effect of water temperature and light intensity on growth rate and toxicity change in Protogonyaulax tamarensis, Mar. Biol., 95, 217, 10.1007/BF00409008

Paerl, 2008, Blooms like it hot, Science, 320, 57, 10.1126/science.1155398

Paerl, 2012, Climate change: links to global expansion of harmful cyanobacteria, Water Res., 46, 1349, 10.1016/j.watres.2011.08.002

Pang, 2017, Effect of CO2 on growth and toxicity of Alexandrium tamarense from the East China Sea, a major producer of paralytic shellfish toxins, Harmful Algae, 68, 240, 10.1016/j.hal.2017.08.008

Pansch, 2014, Habitat traits and food availability determine the response of marine invertebrates to ocean acidification, Glob. Change Biol., 20, 765, 10.1111/gcb.12478

Pazos, 2006, Early detection and intensive monitoring during an unusual toxic bloom of Gymnodinium catenatum advected into the Galician Rias (NW Spain), 12th International Conference on Harmful Algae, Programme and Abstracts, 259

Peng, 2018, Seasonally relevant cool temperatures interact with N chemistry to increase microcystins produced in lab cultures of Microcystis aeruginosa NIES-843, Environ. Sci. Technol., 52, 4127, 10.1021/acs.est.7b06532

Pierce, 2003, Brevetoxin concentrations in marine aerosol: human exposure levels during a Karenia brevis harmful algal bloom, Bull. Environ. Contam. Toxicol., 70, 161, 10.1007/s00128-002-0170-y

Pitcher, 2017, Harmful algal blooms in eastern boundary upwelling systems: a GEOHAB core research project, Oceanography, 30, 22, 10.5670/oceanog.2017.107

Pörtner, 2005, Synergistic effects of temperature extremes, hypoxia, and increases in CO2 on marine animals: From Earth history to global change, J. Geophys. Res., 110, 10.1029/2004JC002561

Pörtner, 2008, Ecosystem effects of ocean acidification in times of ocean warming: a physiologist’s view, Mar. Ecol. Prog. Ser., 373, 203, 10.3354/meps07768

Pörtner, 2010, Oxygen- and capacity-limitation of thermal tolerance: a matrix for integrating climate-related stressor effects in marine ecosystems, J. Exp. Biol., 213, 881, 10.1242/jeb.037523

Pörtner, 2008, Physiology and climate change, Science, 322, 690, 10.1126/science.1163156

Pörtner, 2007, Climate change affects marine fishes through the oxygen limitation of thermal tolerance, Science, 315, 95, 10.1126/science.1135471

Proenca, 2017, Marine and fresh-water harmful algae, Proceedings of the 17th International Conference on Harfmul Algae

Rabalais, 2002, Gulf of Mexico hypoxia, A.K.A. “The Dead Zone.”, Annu. Rev. Ecol. Syst., 33, 235, 10.1146/annurev.ecolsys.33.010802.150513

Ramajo, 2016, Food supply confers calcifiers resistance to ocean acidification, Sci. Rep., 6, 19374, 10.1038/srep19374

Rapala, 1997, Variation of microcystins, cyanobacterial hepatotoxins, in Anabaena spp. as a function of growth stimuli, Appl. Environ. Microbiol., 63, 2206, 10.1128/AEM.63.6.2206-2212.1997

Raven, 2019, Dynamic CO2 and pH levels in coastal, estuarine, and inland waters: theoretical and observed effects of on harmful algal blooms, Harmful Algae, 91

Richlen, 2010, The catastrophic 2008–2009 red tide in the Arabian gulf region, with observations on the identification and phylogeny of the fish-killing dinoflagellate Cochlodinium polykrikoides, Harmful Algae, 9, 163, 10.1016/j.hal.2009.08.013

Rivasseau, 1999, Evaluation of an ELISA Kit for the monitoring of microcystins (myanobacterial toxins) in water and algae environmental samples, Environ. Sci. Tech., 33, 1520, 10.1021/es980460g

Röder, 2012, Effects of salinity, temperature and nutrients on growth, cellular characteristics and yessotoxin production of Protoceratium reticulatum, Harmful Algae, 15, 59, 10.1016/j.hal.2011.11.006

Rountos, 2019, The combined and individual effects of the harmful algae, Alexandrium catenella and Dinophysis acuminata, on the survival, growth, and swimming activity of early life stages of forage fish, Mar. Environ. Res., 10.1016/j.marenvres.2019.04.013

Ryan, 2009, Influences of upwelling and downwelling winds on red tide bloom dynamics in Monterey Bay, California, Cont. Shelf Res., 29, 785, 10.1016/j.csr.2008.11.006

Ryan, 2017, Causality of an extreme harmful algal bloom in Monterey Bay, California, during the 2014-2016 northeast Pacific warm anomaly: extreme HAB Causality in Monterey Bay, Geophys. Res. Lett., 44, 5571, 10.1002/2017GL072637

Sandrini, 2014, Genetic diversity of inorganic carbon uptake systems causes variation in CO2 response of the cyanobacteriumMicrocystis, ISME J., 8, 589, 10.1038/ismej.2013.179

Sherman, 1984, Spawning strategies of fishes in relation to circulation, phytoplankton production, and pulses in zooplankton off the northeastern United States, Mar. Ecol. Prog. Ser., 18, 1, 10.3354/meps018001

Shumway, 1990, A review of the effects of algal blooms on shellfish and aquaculture, J. World Aquac. Soc., 21, 65, 10.1111/j.1749-7345.1990.tb00529.x

Shumway, 2003, Shellfish aquaculture–in praise of sustainable economies and environments, World Aquac., 34, 8

Smayda, 2003, Strategies of marine dinoflagellate survival and some rules of assembly, J. Sea Res., 49, 95, 10.1016/S1385-1101(02)00219-8

Sokolova, 2013, Energy-limited tolerance to stress as a conceptual framework to integrate the effects of multiple stressors, Integr. Comp. Biol., 53, 597, 10.1093/icb/ict028

Sokolova, 2001, Temperature effects on key metabolic enzymes in Littorina saxatilis and L. obtusata from different latitudes and shore levels, Mar. Biol., 139, 113, 10.1007/s002270100557

Sun, 2011, Effects of changing pCO2 and phosphate availability on domoic acid production and physiology of the marine harmful bloom diatom Pseudo-nitzschia multiseries, Limnol. Oceanogr., 56, 829, 10.4319/lo.2011.56.3.0829

Sunda, 2012, Eutrophication induced CO2-acidification of subsurface coastal waters: interactive effects of temperature, salinity, and atmospheric pCO2, Environ. Sci. Technol., 46, 10651, 10.1021/es300626f

Sunda, 2006, Positive feedback and the development and persistence of ecosystem disruptive algal blooms, J. Phycol., 42, 963, 10.1111/j.1529-8817.2006.00261.x

Talmage, 2009, The effects of elevated carbon dioxide concentrations on the metamorphosis, size, and survival of larval hard clams (Mercenaria mercenaria), bay scallops (Argopecten irradians), and eastern oysters (Crassostrea virginica), Limnol. Oceanogr., 54, 2072, 10.4319/lo.2009.54.6.2072

Talmage, 2010, Effects of past, present, and future ocean carbon dioxide concentrations on the growth and survival of larval shellfish, Proc. Natl. Acad. Sci., 107, 17246, 10.1073/pnas.0913804107

Talmage, 2012, Effects of CO2 and the harmful alga Aureococcus anophagefferens on growth and survival of oyster and scallop larvae, Mar. Ecol. Prog. Ser., 464, 121, 10.3354/meps09867

Tang, 2009, Cochlodinium polykrikoides blooms and clonal isolates from the northwest Atlantic coast cause rapid mortality in larvae of multiple bivalve species, Mar. Biol., 156, 2601, 10.1007/s00227-009-1285-z

Tang, 2009, Characterization of the toxicity of Cochlodinium polykrikoides isolates from Northeast US estuaries to finfish and shellfish, Harmful Algae, 8, 454, 10.1016/j.hal.2008.10.001

Tango, 2005, Impacts and potential effects due to Prorocentrum minimum blooms in Chesapeake Bay, Harmful Algae, 4, 525, 10.1016/j.hal.2004.08.014

Tatters, 2012, High CO2 and silicate limitation synergistically increase the toxicity of Pseudo-nitzschia fraudulenta, PLoS One, 7, 10.1371/journal.pone.0032116

Tatters, 2013, High CO2 promotes the production of paralytic shellfish poisoning toxins by Alexandrium catenella from Southern California waters, Harmful Algae, 30, 37, 10.1016/j.hal.2013.08.007

Thomsen, 2013, Food availability outweighs ocean acidification effects in juvenile Mytilus edulis : laboratory and field experiments, Glob. Change Biol., 19, 1017, 10.1111/gcb.12109

Trainer, 2019, Pelagic harmful algal blooms and climate change: Lessons from nature’s experiments with extremes, 91

Trimborn, 2008, Inorganic carbon acquisition in potentially toxic and non-toxic diatoms: the effect of pH-induced changes in seawater carbonate chemistry, Physiol. Plant., 133, 92, 10.1111/j.1399-3054.2007.01038.x

Tyler, 2009, Temporal and spatial dynamics of diel-cycling hypoxia in estuarine tributaries, Estuaries Coasts, 32, 123, 10.1007/s12237-008-9108-x

Valiela, 2009

Van de Waal, 2009, The ecological stoichiometry of toxins produced by harmful cyanobacteria: an experimental test of the carbon-nutrient balance hypothesis: ecological stoichiometry of toxin production, Ecol. Lett., 12, 1326, 10.1111/j.1461-0248.2009.01383.x

Van de Waal, 2011, Reversal in competitive dominance of a toxic versus non-toxic cyanobacterium in response to rising CO2, ISME J., 5, 1438, 10.1038/ismej.2011.28

Van de Waal, 2014, Impact of elevated pCO2 on paralytic shellfish poisoning toxin content and composition in Alexandrium tamarense, Toxicon, 78, 58, 10.1016/j.toxicon.2013.11.011

van der Westhuizen, 1985, Effect of temperature and light on the toxicity and growth of the blue-green alga Microcystis aeruginosa (UV-006), Planta, 163, 55, 10.1007/BF00395897

Vaquer-Sunyer, 2008, Thresholds of hypoxia for marine biodiversity, Proc. Natl. Acad. Sci., 105, 15452, 10.1073/pnas.0803833105

Vaquer-Sunyer, 2011, Temperature effects on oxygen thresholds for hypoxia in marine benthic organisms, Glob. Change Biol., 17, 1788, 10.1111/j.1365-2486.2010.02343.x

Vargas-Yáñez, 2005, Trends and time variability in the northern continental shelf of the western Mediterranean, J. Geophys. Res., 110, 10.1029/2004JC002799

Waldbusser, 2014, Ocean acidification in the coastal zone from an organism’s perspective: multiple system parameters, frequency domains, and habitats, Annu. Rev. Mar. Sceince, 6, 221, 10.1146/annurev-marine-121211-172238

Waldbusser, 2010, Size-dependent pH effect on calcification in post-larval hard clam Mercenaria spp, Mar. Ecol. Prog. Ser., 417, 171, 10.3354/meps08809

Waldbusser, 2011, Biocalcification in the eastern oyster (Crassostrea virginica) in relation to long-term trends in Chesapeake Bay pH, Estuaries Coasts, 34, 221, 10.1007/s12237-010-9307-0

Waldbusser, 2015, Ocean acidification has multiple modes of action on bivalve larvae, PLoS One, 10, 10.1371/journal.pone.0128376

Wallace, 2014, Coastal ocean acidification: the other eutrophication problem, Estuar. Coast. Shelf Sci., 148, 1, 10.1016/j.ecss.2014.05.027

Walls, 2018, Hot and toxic: temperature regulates microcystin release from cyanobacteria, Sci. Total Environ., 610–611, 786, 10.1016/j.scitotenv.2017.08.149

Walther, 2002, Ecological responses to recent climate change, Nature, 416, 389, 10.1038/416389a

Watanabe, 1985, Effects of environmental factors on toxicity of a cyanobacterium (Microcystis aeruginosa) under culture conditions, Appl. Environ. Microbiol., 49, 1342, 10.1128/AEM.49.5.1342-1344.1985

Waycott, 2009, Accelerating loss of seagrasses across the globe threatens coastal ecosystems, Proc. Natl. Acad. Sci., 106, 12377, 10.1073/pnas.0905620106

Wells, 2015, Harmful algal blooms and climate change: learning from the past and present to forecast the future, Harmful Algae, 49, 68, 10.1016/j.hal.2015.07.009

Wikfors, 1995, Experimental and histological studies of four life-history stages of the eastern oyster, Crassostrea virginica, exposed to a cultured strain of the dinoflagellate Prorocentrum minimum, Biol. Bull., 188, 313, 10.2307/1542308

Woodward, 2010, Climate change and freshwater ecosystems: impacts across multiple levels of organization, Philos. Trans. R. Soc. B Biol. Sci., 365, 2093, 10.1098/rstb.2010.0055

Wu, 2017, Histopathological alterations in triangle sail mussel (Hyriopsis cumingii) exposed to toxic cyanobacteria (Microcystis aeruginosa) under hypoxia, Aquaculture, 467, 182, 10.1016/j.aquaculture.2016.05.026

Yu, 2015, Effects of elevated CO2 on dynamics of microcystin-producing and non-microcystin-producing strains during Microcystis blooms, J. Environ. Sci., 27, 251, 10.1016/j.jes.2014.05.047

Zhang, 2011, Studies on the toxic effects of microcystin-LR on the zebrafish (Danio rerio) under different temperatures, J. Appl. Toxicol., 31, 561, 10.1002/jat.1597

Zhang, 2012, Emergence of brown tides caused by Aureococcus anophagefferens Hargraves et Sieburth in China, Harmful Algae, 19, 117, 10.1016/j.hal.2012.06.007

Zhu, 2017, Understanding the blob bloom: warming increases toxicity and abundance of the harmful bloom diatom Pseudo-nitzschia in California coastal waters, Harmful Algae, 67, 36, 10.1016/j.hal.2017.06.004