GnRHs and GnRH receptors

Animal Reproduction Science - Tập 88 Số 1-2 - Trang 5-28 - 2005
Robert P. Millar1
1MRC Human Reproductive Sciences Unit, The Chancellor's Building, 49 Little France Crescent, Edinburgh EH16 4SB, Scotland, UK

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Adams, 2003, Sic novel gonadotropin-releasing hormones are encoded as triplets on each of two genes in the protochordate, Ciona intestinalis, Endocrinology, 144, 1907, 10.1210/en.2002-0216

Alok, 2000, Characterization of a pituitary GnRH-receptor from a perciform fish, Morone saxatilis: functional expression in a fish cell line, Mol. Cell. Endocrinol., 168, 65, 10.1016/S0303-7207(00)00317-8

Arora, 1997, Mutations of the conserved DRS motif in the second intracellular loop of the gonadotropin-releasing hormone receptor affect expression, activation, and internalization, Mol. Endocrinol., 11, 1203, 10.1210/me.11.9.1203

Arora, 1998, Mediation of cyclic AMP signaling by the first intracellular loop of the gonadotropin-releasing hormone receptor, J. Biol. Chem., 273, 25581, 10.1074/jbc.273.40.25581

Baba, 1971, Structure of the porcine LH- and FSH-releasing hormone. II. Confirmation of the proposed structure by conventional sequential analyses, Biochem. Biophys. Res. Commun., 44, 459, 10.1016/0006-291X(71)90623-1

Baldwin, 1993, The probable arrangement of the helices in G protein-coupled receptors, EMBO J., 12, 1693, 10.1002/j.1460-2075.1993.tb05814.x

Ballesteros, 1992, Analysis and refinement of criteria for predicting the structure and relative orientations of transmembranal helical domains, Biophys. J., 62, 107, 10.1016/S0006-3495(92)81794-0

Ballesteros, 1998, Functional microdomains in G-protein-coupled receptors. The conserved arginine-cage motif in the gonadotropin-releasing hormone receptor, J. Biol. Chem., 273, 10445, 10.1074/jbc.273.17.10445

Barbieri, 1992, Clinical applications of GnRH and its analogues, Trends Endocrinol. Metab., 3, 30, 10.1016/1043-2760(92)90089-J

Bertherat, 1998, Gonadotropin-releasing hormone receptor gene mutation: a new cause of hereditary hypogonadism and another mutated G-protein-coupled receptor, Eur. J. Endocrinol., 138, 621, 10.1530/eje.0.1380621

Brooks, 1993, Cloning and sequencing of the sheep pituitary gonadotropin-releasing hormone receptor and changes in expression of its mRNA during the estrous cycle, Mol. Cell. Endocrinol., 94, R23, 10.1016/0303-7207(93)90177-L

Casper, 1991, Clinical uses of gonadotropin-releasing hormone analogues, Can. Med. Assoc. J., 144, 153

Chauvin, 2000, Functional importance of transmembrane Helix 6 Trp279 and Exoloop 3 Val299 of rat gonadotropin-releasing hormone receptor, Mol. Pharmacol., 57, 625, 10.1124/mol.57.3.625

Chi, 1993, Cloning and characterization of the human GnRH receptor, Mol. Cell. Endocrinol., 91, R1, 10.1016/0303-7207(93)90278-R

Chi, 1994, Mutations of the second intracellular loop domain of the GnRH receptor

Conklin, 2000, Cloning and gene expression of a novel human ribonucleoprotein, Biochim. Biophys. Acta, 1492, 465, 10.1016/S0167-4781(00)00090-7

Conn, 1991, Gonadotropin-releasing hormone and its analogues, New Engl. J. Med., 324, 93, 10.1056/NEJM199101103240205

Conn, 1994, Gonadotropin-releasing hormone and its analogs, Ann. Rev. Med., 45, 391, 10.1146/annurev.med.45.1.391

Davidson, 1995, Identification of N-glycosylation sites in the gonadotropin-releasing hormone receptor: role in receptor expression but not ligand binding, Mol. Cell. Endocrinol., 107, 241, 10.1016/0303-7207(94)03449-4

Davidson, 1996, Incorporation of an additional glycosylation site enhances expression of functional human gonadotropin-releasing hormone receptor, Endocrine, 4, 207, 10.1007/BF02738686

Davidson, 1996, Asn102 of the gonadotropin-releasing hormone receptor is a critical determinant of potency for agonists containing C-terminal glycinamide, J. Biol. Chem., 271, 15510, 10.1074/jbc.271.26.15510

Davidson, 1997, Irreversible activation of the gonadotropin-releasing hormone receptor by photoaffinity cross-linking: localization of attachment site to Cys residue in N-terminal segment, Biochemistry, 36, 12881, 10.1021/bi971377t

De Lean, 1980, A ternary complex model explains the agonist-specific binding properties of the adenylate cyclase-coupled beta-adrenergic receptor, J. Biol. Chem., 255, 7108, 10.1016/S0021-9258(20)79672-9

de Roux, 1997, A family with hypogonadotropic hypogonadism and mutations in the gonadotropin-releasing hormone receptor, New Eng. J. Med., 337, 1597, 10.1056/NEJM199711273372205

de Roux, 1999, The same molecular defects of the gonadotropin-releasing hormone-receptor determine a variable degree of hypogonadism in affected kindred, J. Clin. Endocrinol. Metab., 84, 567, 10.1210/jc.84.2.567

Donnelly, 1989, An analysis of the periodicity of conserved residues in sequence alignments of G-protein coupled receptors Implications for the three-dimensional structure, FEBS Lett., 251, 109, 10.1016/0014-5793(89)81438-3

Eidne, 1992, Molecular cloning and characterisation of the rat pituitary gonadotrophn-releasing hormone (GnRH) receptor, Mol. Cell. Endocrinol., 90, R5, 10.1016/0303-7207(92)90116-N

Emons, 1994, The use of luteinizing hormone releasing hormone agonists and antagonists in gynaecological cancers, Hum. Reprod., 9, 1364, 10.1093/oxfordjournals.humrep.a138714

Faurholm, 2001, The genes encoding for the Type II gonadotropin-releasing hormone receptor and the ribonucleoprotein RBM8A overlap in two genomic loci, Genomics, 78, 15, 10.1006/geno.2001.6650

Filicori, 1994, Gonadotrophin-releasing hormone agonists A guide to use and selection [published erratum appears in Drugs Aug;48(2):326]., Drugs, 48, 41, 10.2165/00003495-199448010-00005

Fink, 1988, Gonadotropin secretion and its control, 1349

Flanagan, 1994, Glutamate 301 of the mouse gonadotropin-releasing hormone receptor confers specificity for Arginine8 of mammalian gonadotropin-releasing hormone, J. Biol. Chem., 269, 22636, 10.1016/S0021-9258(17)31693-9

Flanagan, 1997, Advances in understanding gonadotrophin-releasing hormone receptor structure and ligand interactions, Rev. Reprod., 2, 113, 10.1530/ror.0.0020113

Flanagan, 1999, The functional microdomain in transmembrane helices 2 and 7 regulates expression, activation and coupling pathways of the gonadotrophin-releasing hormone receptor, J. Biol. Chem., 274, 28880, 10.1074/jbc.274.41.28880

Fraser, 1993, GnRH analogues for contraception, Br. Med. Bull., 1993, 62, 10.1093/oxfordjournals.bmb.a072606

Fromme, 2001, Role of aspartate 7.32 (302) of the human gonadotropin-releasing hormone receptor in stabilizing a high-affinity ligand conformation, Mol. Pharmacol., 60, 1280, 10.1124/mol.60.6.1280

Guarnieri, 1996, Conformational memories and the exploration of biologically relevant peptide conformations: an illustration for the gonadotropin-releasing hormone, J. Am. Chem. Soc., 118, 5580, 10.1021/ja952745o

Hauser, 1998, Molecular cloning, genomic organization and developmental regulation of a novel receptor from Drosophila melanogaster structurally related to gonadotropin-releasing hormone receptors for vertebrates, Biochem. Biophys. Res. Commun., 249, 822, 10.1006/bbrc.1998.9230

Hoffmann, 2000, Residues within transmembrane helices 2 and 5 of the human gonadotropin-releasing hormone receptor contribute to agonist and antagonist binding, Mol. Endocrinol., 14, 1099, 10.1210/me.14.7.1099

Hsueh, 1985, Gonadotropin-releasing hormone as a paracrine hormone and neurotransmitter in extra-pituitary sites, J. Steroid Biochem., 23, 757, 10.1016/S0022-4731(85)80011-X

Illing, 1993, Comparative sequence analysis and functional characterization of the cloned sheep gonadotropin-releasing hormone receptor reveal differences in primary structure and ligand specificity among mammalian receptors, Biochem. Biophys. Res. Commun., 196, 745, 10.1006/bbrc.1993.2312

Illing, 1999, Two gonadotropin-releasing hormone receptor subtypes with distinct ligand selectivity and differential distribution in brain and pituitary in the goldfish (Carassius autatus), Proc. Natl. Acad. Sci. U.S.A., 96, 2256, 10.1073/pnas.96.5.2526

Jennes, 1994, Gonadotropin-releasing hormone and its receptors in rat brain, Front Neuroendocr., 15, 51, 10.1006/frne.1994.1003

Jimenz-Linan, 1997, Examination of guinea pig luteinizing-hormone-releasing-hormone gene reveals a unique decapeptide and existence of two transcripts in the brain, Endocrinology, 138, 423

Jones, 1987, Chicken II luteinizing hormone-releasing hormone inhibits the M-current of bullfrog sympathetic neurons, Neurosci. Lett., 80, 180, 10.1016/0304-3940(87)90650-1

Kaiser, 1992, Isolation and characterization of cDNAs encoding the rat pituitary gonadotropin-releasing hormone receptor, Biochem. Biophys. Res. Commun., 189, 1645, 10.1016/0006-291X(92)90266-N

Kakar, 1992, Cloning, sequencing, and expression of human gonadotropin releasing hormone (GnRH) receptor, Biochem. Biophys. Res. Commun., 189, 289, 10.1016/0006-291X(92)91556-6

Kakar, 1993, Molecular cloning, sequencing, and characterizing the bovine receptor for gonadotropin releasing hormone (GnRH), Dom. Anim. Endocrinol., 10, 335, 10.1016/0739-7240(93)90037-C

Karten, 1986, Gonadotropin-releasing hormone analog design. Structure-function studies toward the development of agonists and antagonists: rationale and perspective, Endocr. Rev., 7, 44, 10.1210/edrv-7-1-44

Kenakin, 1993

King, 1980, Comparative aspects of luteinizing hormone-releasing hormone structure and function in vertebrate phylogeny, Endocrinology, 106, 707, 10.1210/endo-106-3-707

King, 1992, Evolution of gonadotropin-releasing hormone, Trends Endocrin. Metab., 3, 339, 10.1016/1043-2760(92)90113-F

King, 1995, Evolutionary aspects of gonadotropin-releasing hormone and its receptor, Cell Mol. Neurobiol., 15, 5, 10.1007/BF02069556

King, 1997, Heterogeneity of vertebrate luteinizing hormone-releasing hormone, Science, 206, 67, 10.1126/science.384514

King, 1997, Coordinated evolution of GnRHs and their receptors, 51

King, 2000, Cloning and expression, pharmacological characterization and internalisation kinetics of the pituitary GnRH receptor in a metatherian species of mammal, Gen. Comp. Endocrinol., 117, 439, 10.1006/gcen.1999.7418

Lescheid, 1997, A second form of gonadotropin-releasing hormone (GnRH) with characteristics of chicken GnRH-II is present in the primate brain, Endocrinology, 138, 5618, 10.1210/en.138.12.5618

Lesk, 1992, Homology modelling: inferences from tables of aligned sequences, Curr. Opin. Struct. Biol., 2, 242, 10.1016/0959-440X(92)90153-X

Madigou, 2000, Cloning, tissue distribution, and central expression of the gonadotropin-releasing hormone receptor in the rainbow trout (Oncorhynchus mykiss), Biol. Reprod., 63, 1857, 10.1095/biolreprod63.6.1857

Maliekal, 1997, Solution conformations of gonadotropin releasing hormone (GnRH) and [Gln8] GnRH, S. Afr. J. Chem., 50, 217

Maney, 1997, Central administration of chicken gonadotropin-releasing hormone-II enhances courtship behavior in a female sparrow, Horm. Behav., 32, 11, 10.1006/hbeh.1997.1399

Matsuo, 1971, Structure of the porcine LH- and FSH-releasing hormone. I. The proposed amino acid sequence, Biochem. Biophys. Res. Commun., 43, 1334, 10.1016/S0006-291X(71)80019-0

Maudsley, 2004, Gonadotropin-Releasing Hormone (GnRH) antagonists promote proapoptitic signalling in peripheral reproductive tumor cells by activating a Gxi-coupling state of the type I GnRH receptor, Cancer Research, 64, 7533, 10.1158/0008-5472.CAN-04-1360

Millar, 2002, Gonadotropin releasing hormones and their receptors

Millar, 2003, GnRH II and type II GnRH receptors, Trends Endocrinol. Metab., 14, 35, 10.1016/S1043-2760(02)00016-4

Millar, 1983, Synthesis, luteinizing hormone-releasing activity, and receptor binding of chicken hypothalamic luteinizing hormone-releasing hormone, Endocrinology, 113, 1364, 10.1210/endo-113-4-1364

Millar, 1987, Structural and functional evolution of gonadotropin-releasing hormone, Int. Rev. Cytol., 106, 149, 10.1016/S0074-7696(08)61712-X

Millar, 1988, Evolution of gonadotropin-releasing hormone: multiple usage of a peptide, News Physiol. Sci., 3, 49

Millar, 1987, Gonadotrophin-releasing hormone—diversity of functions and clinical applications, S. Afr. Med. J., 72, 748

Millar, 1989, J. Biol. Chem., 264, 21007, 10.1016/S0021-9258(19)30036-5

Millar, 1997, Plasticity in the structural and functional evolution of GnRH: a peptide for all seasons, 15

Millar, 1999, A novel human GnRH receptor homolog gene: abundant and wide tissue distribution of the antisense transcript, J. Endocrinol., 162, 117, 10.1677/joe.0.1620117

Millar, 2000, Progress towards the development of non-peptide orally-active gonadotropin-releasing hormone (GnRH) antagonists: therapeutic implications, Br. Med. Bull., 56, 761, 10.1258/0007142001903346

Millar, 2001, A novel mammalian receptor for the evolutionarily conserved Type II gonadotropin-releasing hormone, Proc. Natl. Acad. Sci., 98, 9636, 10.1073/pnas.141048498

Millar, 2004, Outside in and inside out signalling: the new concept that selectivity of ligand binding at the GnRH receptor is modulated by the intracellular environment, Endocrinology, 145, 3590, 10.1210/en.2004-0461

Millar, 2004, Gonadotropin-releasing hormone receptors, Endocrine Reviews, 25, 235, 10.1210/er.2003-0002

Milton, 1983, Comparative structure activity studies of mammalian [Arg8]LHRH and chicken [Gln8]LHRH by fluorimetric titration, Biochem. Biophys. Res. Commun., 111, 1082, 10.1016/0006-291X(83)91410-9

Mitchell, 1998, Rhodopsin-family receptors associate with small G proteins to activate phospholipase D, Nature, 392, 411, 10.1038/32937

Miyamoto, 1984, Identification of the second gonadotropin-releasing hormone in chicken hypothalamus: evidence that gonadotropin secretion is probably controlled by two distinct gonadotropin-releasing hormones in avian species, Proc. Natl. Acad. Sci. U.S.A., 81, 3874, 10.1073/pnas.81.12.3874

Moghissi, 1992, Clinical applications of gonadotropin-releasing hormones in reproductive disorders, Endocrinol. Metab. Clin. North. Am., 21, 125, 10.1016/S0889-8529(18)30235-4

Morgan, 2003, A transcriptionally active human Type II gonadotropin releasing hormone receptor gene homologue overlaps two genes in the antisense orientation on chromosome 1q.12, Endocrinology, 144, 423, 10.1210/en.2002-220622

Moss, 1979, Actions of hypothalamic-hypophysiotropic hormones on the brain, Ann. Rev. Physiol., 41, 617, 10.1146/annurev.ph.41.030179.003153

Muske, 1993, Evolution of gonadotropin-releasing hormone (GnRH) neuronal systems, Brain Behav. E, 42, 215, 10.1159/000114156

Myburgh, 1998, Alanine-261 in intracellular loop III of the human gonadotropin-releasing hormone receptor is crucial for G-protein coupling and receptor internalization, Biochem. J., 331, 893, 10.1042/bj3310893

Naor, 1998, Mechanism of GnRH receptor signaling: combinatorial cross-talk of Ca2+ and protein kinase C, Front Neuroendocrinol., 19, 1, 10.1006/frne.1997.0162

Neill, 2001, A gonadotropin-releasing hormone (GnRH) receptor specific for GnRH II in primates, Biochem. Biophys. Res. Commun., 282, 1012, 10.1006/bbrc.2001.4678

Nieschlag, 1992, Hormonal male contraception: a real chance?, 477

Okubo, 2000, A novel form of gonadotropin-releasing hormone in the medaka, Oryzias latipes, Biochem. Biophys. Res. Commun., 16, 298, 10.1006/bbrc.2000.3476

Okubo, 2000, Molecular cloning and tissue specific expression of a gonadotropin-releasing hormone receptor in the Japanese eel, Gen. Comp. Endocrinol., 119, 181, 10.1006/gcen.2000.7511

Okubo, 2001, Identification and characterization of two distinct GnRH receptor subtypes in a teleost, the Medaka Oryzias latipes, Endocrinology, 142, 4729, 10.1210/en.142.11.4729

Palczewski, 2000, Crystal structure of rhodopsin: a G protein-coupled receptor, Science, 289, 739, 10.1126/science.289.5480.739

Perrin, 1993, Molecular and functional ch‘racterization of GnRH receptors cloned from rat pituitary and a mouse pituitary tumor cell line, Biochem. Biophys. Res. Commun., 181, 1139, 10.1006/bbrc.1993.1335

Petry, 2002, Secondary structure of the third extracellular loop responsible for ligand selectivity of a mammalian gonadotropin-releasing hormone receptor, J. Med. Chem., 45, 1026, 10.1021/jm011036k

Powell, 1996, Two new forms of gonadotropin-releasing hormone in a protochordate and the evolutionary implications, Neurobiology, 93, 10461

Reinhart, 1992, Molecular cloning and expression of cDNA encoding the murine gonadotropin-releasing hormone receptor, J. Biol. Chem., 267, 21281, 10.1016/S0021-9258(19)36602-5

Rissman, 1997, Behavioral regulation of gonadotropin-releasing hormone production, Brain. Res. Bull., 44, 459, 10.1016/S0361-9230(97)00226-8

Robison, 2001, Gonadotropin-releasing hormone receptor in the teleost Haplochromis burtoni: structure, location, and function, Endocrinology, 142, 1737, 10.1210/en.142.5.1737

Rodic, 1996, Role of Asp2.61(198) in agonist complexing with the human gonadotropin-releasing hormone receptor, Soc. Neurosci. Abstr., 22, 1302

Samama, 1993, A mutation-induced activated state of the beta 2-adrenergic receptor. Extending the ternary complex model, J. Biol. Chem., 268, 4625, 10.1016/S0021-9258(18)53442-6

Schally, 1971, Isolation and properties of the FSH and LH-releasing hormone, Biochem. Biophys. Res. Commun., 43, 393, 10.1016/0006-291X(71)90766-2

Schertler, 1993, Projection structure of rhodopsin, Nature, 362, 770, 10.1038/362770a0

Sealfon, 1994, The gonadotropin-releasing hormone receptor: structural determinants and regulatory control., vol. 17, 255

Sealfon, 1997, Molecular mechanisms of ligand interaction with the gonadotropin-releasing hormone receptor, Endocr. Rev., 18, 180, 10.1210/er.18.2.180

Seeburg, 1987, The mammalian GnRH gene and its pivotal role in reproduction, Rec. Prog. Horm. Res., 43, 69

Sherwood, 1987, The GnRH family of peptides, Trends Neurosci., 1987, 129, 10.1016/0166-2236(87)90058-0

Sherwood, 1989, The origin of the mammalian form of GnRH in primitive fishes, Fish Physiol. Biochem., 7, 85, 10.1007/BF00004693

Sherwood, 1993, Origin of mammalian gonadotropin-releasing hormones, Endocr. Rev., 14, 241, 10.1210/edrv-14-2-241

Sun, 2000, A chicken gonadotroppin-releasing hormone receptor that confers agonist activity to mammalian antagonists: identification of D-Lys6 in the ligand and extracellular loop two of the receptor as determinants, J. Biol. Chem., 276, 7754, 10.1074/jbc.M009020200

Temple, 2003, An evolutionarily conserved form of gonadotropin-releasing hormone coordinates energy and reproductive behavior, Endocrinology, 144, 13, 10.1210/en.2002-220883

Tensen, 1997, Distinct efficacies for two endogenous ligands on a single cognate gonadoliberin receptor, Eur. J. Biochem., 243, 134, 10.1111/j.1432-1033.1997.0134a.x

Troskie, 1997, Chicken GnRH II-like peptides and a GnRH receptor selective for chicken GnRH II in amphibian sympathetic ganglia, Neuroendocrinology, 65, 396, 10.1159/000127202

Troskie, 1998, Identification of three putative GnRH receptor subtypes in vertebrates, Gen. Comp. Endocrinol., 112, 296, 10.1006/gcen.1998.7156

Troskie, 2000, cDNA cloning, gene expression and ligand selectivity of a novel GnRH receptor expressed in the pituitary and midbrain of Xenopus laevis, Endocrinology, 141, 1764, 10.1210/en.141.5.1764

Tsutsumi, 1992, Cloning and functional expression of a mouse gonadotropin-releasing hormone receptor, Mol. Endocrinol., 6, 1163, 10.1210/me.6.7.1163

Ulloa-Aguirre, 1998, The third intracellular loop of the rat gonadotropin-releasing hormone receptor couples the receptor to Gs- and G(q/11)-mediated signal transduction pathways: evidence from loop fragment transfection in GGH3 cells, Endocrinology, 139, 2472, 10.1210/en.139.5.2472

Wang, 2001, Three distinct types of GnRH receptor characterized in the bullfrog, Proc. Natl. Acad. Sci. U.S.A., 98, 361, 10.1073/pnas.011508498

Wang, 2001, Inhibitory activity of alternative splice variants of the bullfrog GnRH receptor-3 on wild-type receptor signalling, Endocrinology, 142, 4015, 10.1210/en.142.9.4015

Weesner, 1994, Rapid communication: nucleotide sequence of luteinizing hormone- releasing hormone (LHRH) receptor cDNA in the pig pituitary, J. Anim. Sci., 72, 1911, 10.2527/1994.7271911x

White, 1994, A second gene for gonadotropin-releasing hormone: cDNA and expression pattern in the brain, Proc. Natl. Acad. Sci. U.S.A., 91, 1423, 10.1073/pnas.91.4.1423

Yoo, 2000, Molecular cloning, distribution and pharmacological characterisation of a novel gonadotropin-releasing hormone ([Trp8]GnRH) in frog brain, Mol. Cell. Endocrinol., 164, 197, 10.1016/S0303-7207(00)00221-5

Zhou, 1994, A reciprocal mutation supports helix 2 and helix 7 proximity in the gonadotropin-releasing hormone receptor, Mol. Pharmacol., 45, 165

Zhou, 1995, A locus of the gonadotropin-releasing hormone receptor that differentiates agonist and antagonist binding sites, J. Biol. Chem., 270, 18853, 10.1074/jbc.270.32.18853