Four new deciduous species of Hyperacanthus (Rubiaceae: Gardenieae) from western Madagascar: the sofikomba alliance

F. Rakotonasolo1,2, A. P. Davis3
1Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar
2Kew Madagascar Conservation Centre (KMCC), Antananarivo, Madagascar
3Royal Botanic Gardens, Kew, Richmond, UK

Tóm tắt

The genus Hyperacanthus comprises 11 species, nine in Madagascar and two in Africa, although many Madagascan species remain undescribed. All Madagascan species of Hyperacanthus are evergreen trees occurring in humid and semi-humid forest. Four deciduous species occur in the dry forests of northern, western, and southern Madagascar but are unknown to science. These four species, collectively and informally referred to as the sofikomba alliance, are described here: H. decaryi, H. piliformis, H. septentrionale and H. sofikomba. Three of the four species are illustrated and all four are mapped. A key to species, species descriptions, other key taxonomic information, and preliminary conservation assessments are also provided.

Từ khóa


Tài liệu tham khảo

Bachman, S., Moat, J., Hill, A. W., De la Torre J. & Scott, B. (2011). Supporting Red List threat assessments with GeoCAT: geospatial conservation assessment tool. In: V. Smith & L. Penev (eds), e-Infrastructures for data publishing in biodiversity science. ZooKeys 150: 117 – 126. https://doi.org/10.3897/zookeys.150.2109 Bridson, D. M. & Robbrecht, E. (1985). Validation of the African genus Hyperacanthus E. Mey. (Rubiaceae tribe Gardenieae). Kew Bull. 40: 273 – 286. https://doi.org/10.2307/4108261 Davis, A. P. & Rakotonasolo, F. (2003). Gardenia and related Genera (Tribe Gardenieae Subtribe Gardeniinae). Pp. 441 – 442. In: S. M. Benstead & J. P. Goodman (eds), The Natural History of Madagascar. University of Chicago Press. Hallé, N. (1967). Étude biologique et morphologique de la tribu des Gardéniées (Rubiaceae). Mém. ORSTOM 22: 1 – 146. Holmgren, P. K., Holmgren, N. H. & Barnett, L. C. (1990). Index Herbariorum. Part 1: The Herbaria of the World, 8th ed. Regnum Veg. New York Botanical Garden. International Union for Conservation of Nature (IUCN) (2012). IUCN Red List Categories and Criteria. Version 3.1. Second edition https://portals.iucn.org/library/sites/library/files/documents/RL-2001-001-2nd.pdf. [Accessed December 2022] Kainulainen, K. & Bremer, B. (2014). Phylogeny of Euclinia and allied genera of Gardenieae (Rubiaceae), and description of Melanoxerus, an endemic genus of Madagascar. Taxon 63: 819 – 830. https://doi.org/10.12705/634.2 Leroy, J.-F. (1974). Recherches sur les Rubiacées de Madagascar, les genres Mantalania et Pseudomantalania (Gardeniées). Adansonia, sér. 2, 14: 29 – 52. https://www.biodiversitylibrary.org/item/281085#page/31/mode/1up Punt, W., Hoen, P. P., Blackmore, S., Nilsson, S. & Le Thomas, A. (2007). Glossary of pollen and spore terminology. Rev. Palaeobot. Palynol. 143: 1 – 81. https://doi.org/10.1016/j.revpalbo.2006.06.008 Rakotonasolo, F. (2007). Etude de la Tribu Gardenieae (Rubiaceae) Malgache, En Particulier le Genre Hyperacanthus E. Mey Ex Bridson. PhD Thesis. l’Université d’Antananarivo. ____ & Davis, A. P. (2001). Hyperacanthus ambovombensis (Rubiaceae): a new species and new generic record for Madagascar. Kew Bull. 56: 945 – 953. https://doi.org/10.2307/4119306 ____ & ____ (2002). Notes on the genus Hyperacanthus (Rubiaceae) including the description of a new species from Madagascar: H. grevei. Kew Bull. 57: 955 – 962. https://doi.org/10.2307/4115725 ____ & ____ (2004). A new and endangered species of Hyperacanthus (Rubiaceae) from SE Madagascar: H. mandenensis. Novon 14: 327 – 331. https://www.jstor.org/stable/3393294 ____ & ____ (2006). Six species of Madagascan Genipa transferred to Hyperacanthus (Rubiaceae-Gardenieae) and new data on general morphology, placentation and ovary structure in Hyperacanthus. Taxon 55: 387 – 396. https://doi.org/10.2307/25065586 Rakotonirina, N., Rakouth, B. & Davis. A. P. (2012). A taxonomic revision of Madagascan Gardenia (Rubiaceae, Gardenieae). Nord. J. Bot. 30: 712 – 728. https://doi.org/10.1111/j.1756-1051.2012.01155.x Reitsma, T. (1969). Size modifications of recent pollen grains under different treatments. Rev. Palaeobot. Palynol. 9: 175 – 202. http://dx.doi.org/https://doi.org/10.1016/0034-6667(69)90003-7 Robbrecht, E. & Puff, C. (1986). A survey of the Gardenieae and related tribes (Rubiaceae). Bot. Jahrb. Syst. 108: 63 – 137. Schols, P., Dessein, S. & Smets, E. (2001). iMap version 2.0. Available from http://www.biovolution.com. [Accessed June 2022] ____, ____, D’hondt, C., Huysmans, S. & Smets, E. (2002). CARNOY: a new digital measurement tool for palynology. Grana 41: 124 – 126. https://doi.org/10.1080/001731302760156936 Sonké, B. (1999). Oxyanthus (Rubiaceae-Gardenieae-Gardeniinae) en Afrique centrale: étude systématique. Opera Bot. Belg. 8: 1 – 106. Thiers, B. M. (2020). Index Herbariorum: A Global Directory of Public Herbaria and Associated Staff. New York Botanical Garden's Virtual Herbarium. http://sweetgum.nybg.org/science/ih/. [Accessed June 2022]