Continuum or zonation? Altitudinal gradients in the forest vegetation of Mt. Kilimanjaro

Barkman J.J. (1973). Synusial approaches to classification. In: Whittaker R.H. (ed) Ordination and classification of communities. Handbook of vegetation science V. Junk, The Hague, pp. 437–491

Beals E.W. (1969). Vegetational change along altitudinal gradients. Science165: 981–985

Beentje H.J. (1994). Kenya Trees, Shrubs and Lianas. National Museums of Kenya, Nairobi

Boughey A.S. 1955a. The vegetation of the mountains of Biafra. In: Proc. Linn. Soc. Lond. Session 165, 1952–1953. Pt. 2, pp. 144–150.

Boughey A.S. (1955b). The nomenclature of the vegetation zones on the mountains of tropical Africa. Webbia 11: 413–423

Braun-Blanquet J. (1964). Pflanzensoziologie. Springer, Wien

Bray J.R. and Curtis J.T. (1957). An ordination of the upland forest communities of southern Wisconsin. Ecol. Monographs 27: 325–349

Bruijnzeel L.A. and Veeneklaas E.J. (1998). Climatic conditions and tropical montane forest productivity: The fog has not lifted yet. Ecology 79: 3–9

Clutton-Brock T.H. and Gillett J.B. (1979). A survey of forest composition in the Gombe National Park, Tanzania. Afr. J. Ecol. 17: 131–158

Colwell R.K. and Hurtt G.C. (1994). Nonbiological gradients in species richness and a spurious Rapoport effect. Am. Nat. 144: 570–595

Curtis J.T. (1959). The vegetation of Wisconsin. An ordination of plant communities. Univ. of Wisconsin Press, Madison

Daubenmire R. (1966). Vegetation: Identification of typal communities. Science 151: 291–298

Frahm J.-P. (1994). Scientific results of the BRYOTROP Expedition to Zaire and Rwanda. 2. The altitudinal zonation of the bryophytes on Mt. Kahuzi, Zaire. Tropical Bryology 9: 153–167

Friis I. and Lawesson J.E. (1993). Altitudinal zonation in the forest tree flora of Northeast tropical Africa. Opera Botanica 121: 125–127

Friis I. (1992). Forest and forest trees of northeast tropical Africa – their natural habitats and distribution patterns in Ethiopia, Djibouti and Somalia. Kew Bulletin Additional Series XV. Her Majesty’s Stationery Office, London

*FTEA (1952). Flora of Tropical East Africa. Royal Botanic Garden, Kew

Gentry A.H. and Dodson C.H. (1987a). Contribution of nontrees to species richness of a tropical rain forest. Biotropica 19: 149–156

Gentry A.H. 1995. Patterns of diversity and floristic composition in neotropical montane forests. In: Churchill S.P., Balslev H., Forero E. and Lurteyn J.L. (eds), Biodiversity and Conservation of Neotropical Montane Forests. Proceedings of the Neotropical Montane Forest Biodiversity and Conservation Symposium. The New York Botanical Garden, pp. 103–126.

Givnish T.J. (1999). On the causes of gradients in tropical tree diversity. J. Ecol. 87: 193–210

Grubb P.J. and Whitmore T.C. (1966). A comparison of montane and lowland rain forest in Ecuador. II. The climate and its effects on the distribution and physiognomy of the forests. J. Ecol. 54: 303–333

Grubb P.J. (1971). Interpretation of the ‘Massenerhebung’ effect on tropical Mountains. Nature 229: 44–45

Grubb P.J. (1977). Control of forest growth and distribution on wet tropical mountains: with special reference to mineral nutrition. Ann. Rev. Ecol. Syst. 8: 83–107

Hall J.B. and Swaine M.D. (1976). Classification and ecology of closed-canopy forest in Ghana. J. Ecol. 64: 913–951

Hamilton A.C. (1975). A quantitative analysis of altitudinal zonation in Uganda forests. Vegetatio 30: 99–106

Hamilton A.C. and Perrott R.A. (1981). A study of altitudinal zonation in the montane forest belt of Mt. Elgon, Kenya/Uganda. Vegetatio 45: 107–125

Hamilton A.C., Ruffo C.K., Mwasha I.V., Mmari C. and Lovett J.C. 1989. A survey of forest types on the East Usambaras using the variable – area tree plot method. In: Hamilton A.C. and Bensted-Smith R. (eds), Forest Conservation in the East Usambara Mountains Tanzania. The IUCN Tropical Forest Programme, pp. 213–225.

Hastenrath S.L. (1967). Rainfall Distribution and Regime in Central America. Archiv für Meteorologie, Geophysik und Bioklimatologie, Ser. B: Allgemeine und biologische Klimatologie 15(3): 201–241

Hastenrath S. (1973). Observations on the periglacial morphology of Mts. Kenya and Kilimanjaro, East Africa. Z. Geomorph. N. F. 16: 161–179

Hedberg O. (1951). Vegetation belts of the East African mountains. Svensk Bot. Tidskrift. 45: 140–202

Hemp A. and Beck E. (2001). Erica excelsa as a component of Mt. Kilimanjaro’s forests. Phytocoenologia 31: 449–475

Hemp A. (2001a). Ecology of the pteridophytes on the southern slopes of Mt. Kilimanjaro. Part II: Habitat selection. Plant Biology 3: 493–523

Hemp A. 2001b. Life form and strategies of forest ferns on Mt. Kilimanjaro. In: Gottsberger G. and Liede S. (eds), Life Forms and Dynamics in Tropical Forests. Disserationes Botanicae 346, pp. 95–130.

Hemp A. (2002a) Ecology of the pteridophytes on the southern slopes of Mt. Kilimanjaro. Part I: Altitudinal distribution. Plant Ecology 159: 211–239

Hemp A. 2002b. Fränkische Steppenheide, tropische Bergregenwälder – zwei gegensätzliche Florenregionen, eine gemeinsame Klassifikationsmethode? Habilitationsschrift, Universität Bayreuth.

Hemp A. 2005a. The banana forests of Kilimanjaro: biodiversity and conservation of the Chagga home gardens. Biodiv. Conserv. http://dx.doi.org/10.1007/s10531-004-8230-8.

Hemp A. (2005b). Climate change-driven forest fires marginalize the impact of ice cap wasting on Kilimanjaro. Global Change Biology 11: 1013–1023

Hemp A. (2005c) The impact of fire on diversity, structure and composition of Mt Kilimanjaro’s vegetation. In: Spehn E., Liberman M., Körner C. (eds) Land use changes and mountain biodiversity. CRC Press LLC, Boca Raton FL, USA (in press)

Hemp A. (a). Altitudinal zonation and diversity patterns of the forests of Mt. Kilimanjaro. In: Bruijnzeel S. and Juvik J. (eds), Mountains in the Mist: Science for Conserving and Managing Tropical Montane Cloud Forests (In press).

Hemp A. (b). Ecology and altitudinal zonation of pteridophytes on Mt. Kilimanjaro. Proceedings of the XVIIth AETFAT Congress 21–26 September 2003 (In press).

Hemp A. (c). Kilimanjaro and its missing bamboo zone. Mountain Research and Development. (In press)

Hemp C. (2001). Aerotegmina, a new genus of African Listroscelidinae (Orthoptera: Tettigoniidae, Listroscelidinae, Hexacentrini). Journal of Orthoptera Research 10: 125–132

Hemp C. (2005). The Chagga home gardens – relict areas for endemic Saltatoria species (Insecta: Orthoptera) on Mt. Kilimanjaro. Biological Conservation 125: 203–210

Holdridge L.R. (1967). Life Zone Ecology. Tropical Science Center San José, Costa Rica

Jacobsen W.B.G. and Jacobsen N.H.G. (1989). Comparision of the pteridophyte floras of Southern and Eastern Africa, with special reference to high-altitude species. Bull. Jard. Bot. Nat. Belg. 59: 261–317

Kessler M. and Bach K. (1999). Using indicator families for vegetation classification in species-rich Neotropical forests. Phytocoenologia 29: 485–502

Kessler M. (2000). Altitudinal zonation of Andean cryptogam communities. J. Biogeog. 27: 275–282

Kitayama K. (1992). An altitudinal transect study of the vegetation on Mt. Kinabalu, Borneo. Vegetatio 102: 149–171

Kitayama K. and Mueller-Dombois D. (1992). Vegetation of the wet windward slope of Haleakala, Maui, Hawaii. Pacific Science 46: 197–220

Kress W.J. (1986). The systematic distribution of vascular epiphytes: an update. Selbyana 9: 2–22

Kürschner H. (1990). Höhengliederung von epiphytischen Laub- und Lebermoosen in Nord-Borneo (Mt. Kinabalu). Nova Hedwigia 51: 77–86

Lambrechts C., Woodley B., Hemp A., Hemp C. and Nnyiti P. (2002). Aerial survey of the threats to Mt. Kilimanjaro forests. UNDP, Dar es Salaam

Lauer W. 1976. Klimatische Grundzüge der Höhenstufung tropischer Gebirge. Tagungsber. wiss. Abh. 40. dt. Geographentag Innsbruck, Wiesbaden, pp. 76–90.

Lauer W., Rafiqpoor M.D. and Frankenberg P. (1996). Die Klimate der Erde. Eine Klassifikation auf ökophysiologischer Grundlage der realen Vegetation. Erdkunde 50: 275–300

Legendre P. and Legendre L. (1998). Numerical Ecology. 2 nd English Edition. Developments in Environmental Modelling 20. Elsevier, Amsterdam

Legendre P. and Vaudor A. 1991. The R Package: Multidimensional Analysis, Spatial Analysis. Département de sciences biologiques, Université de Montréal.

Legendre P., Dallot S. and Legendre L. (1985). Succession of species within a community: Chronological clustering, with applications to marine and freshwater zooplankton. Am. Nat. 125: 257–288

Lieberman D., Lieberman M., Peralta R. and Hartshorn G.S. (1996). Tropical forest structure and composition on a large-scale altitudinal gradient in Costa Rica. J. Ecol. 84: 137–152

Lind E.M. and Morrison M.E.S. (1974). East African Vegetation. Longman, London

Lovett J.C. (1996). Elevational and latitudinal changes in tree associations and diversity in the Eastern Arc mountains of Tanzania. J. Trop. Ecol. 12: 629–650

Lovett J.C. (1998). Continuous change in Tanzanian moist forest tree communities with elevation. J. Trop. Ecol. 14: 719–722

McIntosh R.P. (1967). The continuum concept of vegetation. Botan. Rev. 33: 130–187

Moravec J. (1989). Influences of the individualistic concept of vegetation on syntaxonomy. Vegetatio 81: 29–39

Moreau R.E. (1935). A synecological study of Usambara, Tanganyika Territory, with particular reference to birds. J. Ecol. 23: 1–43

Mueller-Dombois D. and Ellenberg H. (1974). Aims and methods of vegetation ecology. Wiley and Sons, New York

Müller M.J. (1983). Handbuch ausgewählter Klimastationen der Erde. Forschungsstelle Bodenerosion der Universität Trier Mertesdorf (Ruwertal), Trier

Pócs T. (1976) Bioclimatic studies in the Uluguru Mountains (Tanzania, East Africa). II. Correlations between orography, climate and vegetation. Acta Bot. Acad. Scient. Hungaricae 22(1/2): 163–183

Pócs T. 1994. The altitudinal distribution of Kilimanjaro bryophytes. In: Seyani J.H. and Chikuni A.C. (eds), Proceedings of XIII Plenary Meeting AETFAT. Malawi, pp. 797–812.

Richards P.W. (1996). The tropical rain forest. Cambridge Univ. Press, Cambridge

Richter M. (1996). Klimatologische und pflanzenmorphologische Vertikalgradienten in Hochgebirgen. Erdkunde 50: 205–237

Thompson L.G., Mosley-Thompson E., Davis M.E., Henderson K.A., Brecher H.H., Zagorodnov V.S., Mashiotta T.A., Lin P.-N., Mikhalenko V.N., Hardy D.R. and Beer J. (2002). Kilimanjaro ice core records: Evidence of holocene climate change in tropical Africa. Science 298: 589–593

Tuomisto H. and Ruokolainen K. (1993). Distribution of Pteridophyta and Melastomataceae along an edaphic gradient in an Amazonian rain forest. J. Veg. Sci. 4: 25–34

Tuomisto H., Ruokolainen K., Kalliola R. Linna A., Danjoy W. and Rodriguez Z. (1995). Dissecting Amazonian Biodiversity. Science 269: 63–66

Tuomisto H., Ruokolainen K., Aguilar M. and Sarmiento A. (2003). Floristic patterns along a 43-km long transect in an Amazonian rain forest. J. Ecol. 91: 743–756

Van Steenis C.G.G.J. (1984). Floristic altitudinal zones in Malaysia. Bot. J. Linn. Soc. 89: 289–292

Vázquez G.J.A. and Givnish T.J. (1998). Altitudinal gradients in tropical forest composition, structure, and diversity in the Sierra de Manantlán. J. Ecol. 86: 999–1020

Walter H., Harnickell E. and Mueller-Dombois D. (1975). Climate-diagram maps of the individual continents and the ecological climatic regions of the earth. Springer, Berlin

Walter H. and Lieth H. 1967. Klimadiagramm–Weltatlas. Jena.

Whittaker R.H. (1962). Classification of natural communities. Bot. Rev. 28: 1–239

Whittaker R.H. (1967). Gradient analysis of vegetation. Biol. Rev. 49: 207–264

Woldu Z., Feoli E. and Nigatu L. (1989). Partitioning an elevation gradient of vegetation from southeastern Ethiopia by probabilistic methods. Vegetatio 81: 189–198