1Department of Biogeography and Global Change (BGC-MNCN-CSIC), National Museum of Natural Sciences, CSIC, c/Serrano 155bis, E-28006, Madrid, Spain
2Laboratorio Ecotono, INIBIOMA-CONICET, Universidad Nacional del Comahue, Black River, Argentina
3Museu de Ciències Naturals de Granollers (MCNG), E-08402 Granollers, Barcelona, Spain
4Department of Biology, Unit of Zoology, Faculty of Sciences, Universidad Autónoma de Madrid, C/Darwin 2, Campus Universitario de Cantoblanco, 28049 Madrid, Spain