A large-scale simulation of the piriform cortex by a cell automaton-based network model

IEEE Transactions on Biomedical Engineering - Tập 49 Số 9 - Trang 921-935 - 2002
E.T. Claverol1, A.D. Brown2, J.E. Chad3
1Department of Biology, California Institute of Technology, Pasadena, CA, USA
2Electronic System Design Group, Electronics Department, Southampton University, Hampshire, UK
3Centre for Neurosciences, Southampton University, Hampshire, UK

Tóm tắt

An event-driven framework is used to construct a physiologically motivated large-scale model of the piriform cortex containing in the order of 10/sup 5/ neuron-like computing units. This approach is based on a hierarchically defined highly abstract neuron model consisting of finite-state machines. It provides computational efficiency while incorporating components which have identifiable counterparts in the neurophysiological domain. The network model incorporates four neuron types, and glutamatergic excitatory and GABA/sub A/ and GABA/sub B/ inhibitory synapses. The spatio-temporal patterns of cortical activity and the temporal and spectral characteristics of simulated electroencephalograms (EEGs) are studied. In line with previous experimental and compartmental work, 1) shock stimuli elicit EEG profiles with either isolated peaks or damped oscillations, the response type being determined by the intensity of the stimuli, and 2) temporally unpatterned input generates EEG oscillations supported by model-wide waves of excitation.

Từ khóa

#Brain modeling #Large-scale systems #Neurons #Computational modeling #Computational efficiency #Electroencephalography #Biological system modeling #Aggregates #Artificial neural networks #Automata

Tài liệu tham khảo

10.1007/BF00203003 10.1126/science.2315702 10.1088/0954-898X/8/2/004 barkai, 1994, modulation of the input/output function of rat piriform cortex pyramidal cells, J Neurophysiol, 72, 644, 10.1152/jn.1994.72.2.644 wilson, 1988, a computer simulation of olfactory cortex with functional implications for storage and retrieval of olfactory information, Neural Information Processing Systems, 114 ketchum, 1993, synaptic events that generate fast oscillations in piriform cortex, J Neurosci, 13, 3980, 10.1523/JNEUROSCI.13-09-03980.1993 wilson, 1992, cortical oscillations and temporal interactions in a computer simulation of piriform cortex, J Neurophysiol, 67, 981, 10.1152/jn.1992.67.4.981 barkai, 1994, modulation of associative memory function in a biophysical simulation of rat piriform cortex, J Neurophysiol, 72, 659, 10.1152/jn.1994.72.2.659 harberly, 1989, olfactory cortex: model circuit for study of associative memory?, TINS, 12, 258 10.1016/S0925-2312(98)00148-9 10.1016/S0925-2312(01)00629-4 10.1109/TSE.1987.233203 10.1016/S0006-3495(62)86953-7 klee, 1977, computed potentials of cortically arranged populations of neurons, J Neurophysiol, 40, 647, 10.1152/jn.1977.40.3.647 press, 1992, Numerical Recipes in C 10.1016/0006-8993(78)90891-0 10.1023/A:1008893429695 10.1038/scientificamerican0291-78 10.1113/jphysiol.1952.sp004764 10.1162/neco.1997.9.6.1179 10.1007/978-1-4612-1634-6 hasselmo, 1995, cholinergic modulation of activity-dependent synaptic plasticity in the piriform cortex and associative memory function in a network biophysical simulation, J Neurosci, 15, 6592, 10.1523/JNEUROSCI.15-10-06592.1995 10.1007/BF02802549 10.1088/0954-898X/2/2/002 freeman, 1992, Neural Networks Algorithms Applications and Programming Techniques koch, 1998, Methods in Neural Modeling From Ions to Networks 10.1002/cne.902160107 freeman, 1968, relations between unit activity and evoked potentials in prepyriform cortex in cats, J Neurophysiol, 31, 337, 10.1152/jn.1968.31.3.337 10.1007/s004220050482 10.1016/0013-4694(84)90128-7 rall, 1977, core conductor theory and cable properties of neurons, Handbook of physiology, 39 nunez, 1981, Electric Fields of the Brain The Neurophysics of EEG ketchum, 1993, membrane currents evoked by afferent fiber stimulation in rat piriform cortex i. current source-density analysis, J Neurophys, 69, 248, 10.1152/jn.1993.69.1.248 10.1103/PhysRevE.51.738