Echinoids from the Belpberg Beds (Obere Meeresmolasse, Middle Burdigalian) in the area of Bern (Switzerland)

The present paper deals with the echinoid fauna of the Belpberg Beds, a lithostratigraphic unit in the Upper Marine Molasse cropping out southeast of Bern (Switzerland). In this unit, echinoids, in contrast to mollusks, are rare and of low diversity. The scarcity of echinoids is related to the environmental setting dominated by monotonous siliciclastic deposits and strong diagenetic alteration. The echinoid fauna consists of three species Psammechinus dubius dubius, Echinocardium deikei and Hemipatagus ocellatus? Psammechinus dubius dubius and H. ocellatus typically occur in Burdigalian sediments in the Paratethys and help to confirm the dating obtained from the microfossils and molluscs of the Belpberg Beds. Furthermore, the Belpberg echinoid fauna is especially relevant for the palaeobiogeography with species common to both the Rhône Basin and the Central Paratethyan faunas.