1Institute of Healthy Ageing and Department of Genetics, Evolution and Environment, University College London, London WC1E 6BT, UK
2Max Planck Institute for Biology of Ageing, Köln 50931, Germany
3Berlin Institute of Health, Berlin 10117, Germany
4Behavior and Metabolism Laboratory, Champalimaud Centre for the Unknown, Lisbon 1400-038, Portugal
5UCL Ear Institute, University College London, London WC1X 8EE, UK
6Institute of Animal Genetics and Breeding, Sichuan Agricultural University, Chengdu 611130, China
7Charles Perkins Centre, School of Life and Environmental Sciences, University of Sydney, Sydney 2050, Australia