1College of Life Sciences, Key Laboratory of Conservation Biology for Endangered Wildlife, Ministry of Education, Zhejiang University, Zijinggang Campus, Hangzhou, 310058, P. R. China
2State Key Laboratory of Vegetation and Environmental Change, Institute of Botany, Chinese Academy of Sciences, Beijing, 100093, China
3Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, Tubney House, Tubney, Abingdon, OXON OX13 5QL, United Kingdom
4Wildlife Conservation, Department of Animal Health Technology, Yamazaki College of Animal Health Technology, Japan