1Department of Natural Resources and Environmental Management, University of Hawaii at Manoa, 1910 East-West Rd., Sherman 101, Honolulu, HI 96822, USA
2Joint Institute for Marine and Atmospheric Research, University of Hawaii at Manoa, 1000 Pope Rd., Marine Science Building 312, Honolulu, HI 96822, USA
3Australian Research Council Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, QLD 4811, Australia
4Pacific Islands Fisheries Science Center, NOAA National Marine Fisheries Service, 1845 Wasp Blvd, Building 176, Honolulu, HI 96818, USA